OF CYSTOCARPS IN CALLOPHYLLIS LACINIATA. 207 
belong to the carpogonial branch, or whether some of them may 
not arise from other neighbouring branches. 
The carpogone with its trichogyne is borne upon one of the 
smaller cells, which again lies alongside of a large auxiliary cell. 
The trichogyne has often, though not always, the little coil 
near its base, so characteristic of Polyides and some other 
Floridex. It forces its way between the cortical cells, and 
finally protrudes, sometimes to the extent of a third of its 
length. 
At alater stage, presumably after fertilization, fusion takes 
place between the carpogone and the auxiliary cells. 
At this stage the procarp is an irregularly-shaped mass of 
fused cells with projections on all sides. After staining with 
methyl-green, one of the procarps was shown to contain a con- 
siderable number of nuclei, the result evidently of the fusion of 
the different cells and of subsequent divisions of these nuclei. 
The remains of the trichogyne can be traced in such procarps, 
but the basal portion of the carpogone can no longer be distin- 
guished, having evidently become confluent with the next 
auxiliary cell. 
Each cystocarp contains a varying number of these procarps ; 
sometimes they are separated from each other by a mass of tissue, 
sometimes they are crowded together in immediate contact. They 
remain, however, separate and distinct ; in no case was there any 
proof of the existence of fertilizing filaments, such as occur in 
Polyides, Dudresnaya, &e., nor is there any evidence that fusion 
takes place between the cells of different procarpial groups. 
The procarp next divides up by walls parallel or at right 
angles to each other into a mass of large cells. These cells again 
divide by walls in all directions (fig. 4). 
At this stage the primary and secondary divisions can be 
distinctly traced, and the whole procarp breaks up; there does 
not seem to be any sterile tissue left. 
From the ultimate divisions we get the spores. These, as has 
been already mentioned, lie in groups, each of which is the pro- 
duct of a single procarp. The groups may be quite isolated and 
completely surrounded by the compressed thallus-tissue, or they 
may lie so closely together that several groups seem merged into 
one. This is easily intelligible, as at an early stage two or more 
procarps are often found in close contact. 
