IN PLANTS HAVING IRREGULAR COROLLAS., 393 
Nevertheless, in the case of flowers presenting this modification, 
it was often found that the tissues were so bent from their 
original position that the division between the two spurs came 
to lie in or near the median plane of symmetry of the flower. 
To such a degree did this occur, that it was only by tracing up 
the bundles in the walls of the spurs that it could be determined 
from which petal they were developed. In several cases, how- 
ever, the original symmetry was still shown by their position, 
No. 2. Flower having one posterior (purple) petal and four 
anterior (yellow) petals (Plate L. figs. 5 & 6).—This form is very 
common throughout our region, and occurs on a large pro- 
portion of the plants, though perhaps not quite so commonly 
as the pelorie type. Such flowers were never seen with less 
than two spurs, but occasionally they have three (Plate L. 
fig. 6). In all cases examined, the five stamens were all de- 
veloped. 
The calyx of these flowers was placed in such a position that 
the median vertical plane of the flower fell between two pos- 
terior sepals. Hence, though we are not able to state what the 
relations to the floral axis have been developmentally, yet the 
appearances decidedly suggest that this flower may be compared 
with the normal flower by imagining that the median plane of 
the zygomorphy has been deflected so that it falls upon a pos- 
terior petal instead of on an anterior one. What the circum- 
stances may be which lead to this alteration of the plane about 
which the symmetry of the flower is formed, we can offer no 
suggestion, but we shall offer another case of a similar pheno- 
menon in describing the variations of Gladiolus (vide p. 400). 
Fig. 4 represents a flower which is in a manner intermediate 
between Nos. 1 and 2. For in it one of the posterior petals: is 
partially coloured yellow, and is to some extent united to the 
adjacent petal of the anterior lip. Cases of this kind, in which 
a petal does not stand truly with either the anterior or posterior 
lip, are decidedly uncommon. Such intermediate flowers always 
had ¢wo spurs and five stamens. 
A form was frequently found (not figured) which conformed 
exactly with that shown in fig. 5, excepting for the fact that the 
deep cleft which separates the purple petal from the yellow ones 
was less deeply cut on the one side than on the other. These 
flowers had at least two spurs and five stamens. Inasmuch as 
the distinction between the two lips is in such cases partially 
2x42 
