410 MR. W. AND MISS A. BATESON ON FLORAL VARIATIONS 
ll. ABNORMAL FLOWERS, NOT PELORIC. 
3. Flower with cruciform, 4-petalled corolla, having one pos- 
terior partially unmarked petal.—Only two flowers of this type 
were seen (Pl. L. fig. 24). In one the calyx was in five parts, 
like that of the normal flower; the corolla was divided into four 
almost equal lobes, arranged in a cruciform manner. Three of 
the lobes were marked with dark blue, whilst one of them was 
less completely marked than the others. The four stamens were 
arranged in the same manner as that described above for other 
pelorie flowers. Another flower having the same type of corolla 
had only two stamens as in normal flowers. The types 2 and 3 
are especially interesting as showing how bilabiate, pentamerous 
flowers can abruptly assume a tetramerous and cruciform cha- 
racter, either losing their zygomorphic character altogether as in 
type 2, or retaining it in part as in type 3. 
4. Flowers with 4-petalled, bilabiate corolla. Ot this type we 
have had three specimens, two on S. Revii, the other on 
Streptocarpus sp. The corolla in these flowers consisted of two 
upper smaller petals without markings, and two lower, larger 
petals with markings. In both these flowers there was one 
fertile stamen only, which was situated in the median, ventral 
line, between the two marked petals (Pl. LI. fig. 3), and two 
rudimentary stamens occurred laterally. Of the nature of the 
calyx in these flowers we have no record. 
It is difficult to see how this type can have arisen unless by a 
discontinuous process ; for by any continuous process the median 
anterior petal (Pl. LI. fig. 1) must have become gradually 
reduced, and the two stamens (S', S?) on either side of it have 
fused into the single median one which is found in this type 
(Pl. LI. fig. 3). In the specimens examined there was no trace 
of any anterior odd petal or of any second stamen*. 
5. Flowers with 5-petalled bilaterally symmetrical corolla ; 
3 petals being posterior and 2 anterior.— The two flowers of this 
* An abnormal arrangement of corolla and stamens, of an exactly analogous 
nature, we have observed in a species of ZEschyaanthus (longiflora ?). This 
flower has usually a corolla of the same pattern as that of Streptocarpus, and 
two pairs of perfect stamens, occupying a position similar to those of the two 
perfect and two imperfect stamens in Streptocarpus. In the abnormal flower 
the arrangement was still perfectly bilaterally symmetrical, the corolla being 
bilabiate with two petals in each lip, and there were three perfect stamens—one 
long, unpaired, median and anterior; and two shorter, paired and lateral. 
