MR. G. BEKTHAM OX EUPHOBBTACE^. 237 



much enlarged foliaceous fruitiug calyx, with external glands or 

 small accessory lobes alternating with the sepals at their base. 

 Mueller makes a subtribe of this species, characterized by the 

 involucrate female flower ; but there must have been some con- 

 fusion in his idea of this involucre. In the subtribal character 

 he says of the female flower, " Involucrum calyciforme uniflorum," 

 in which he seems to have had in view the large calyx, which at 

 first might be readily taken for an involucre. In the description 

 of the species he corrected this into " Calycis fceminei lacinise 

 . . . . involucrum simulantes," and, abandoning the term invo- 

 lucre, describes the female calyx as " calyculatus .... Calyculi 

 lacinise .... valide biglandulosae, calyce breviores .... sub- 

 deciduae" — a description not quite correct. These appendages 

 are not segments of a continuous involucre, but perfectly detached 

 and distinct from each other, and consist of one rather large gland 

 (never two in our specimens), usually bearing a pubescent, thick- 

 ish, somewhat lanceolate dorsal appendage, sometimes one fourth 

 of the length of the flowering calyx, not one tenth of the fully 

 developed fruiting calyx,, and sometimes the gland is almost bare. 

 These appendages appear to me, therefore, to be perfectly homo- 

 logous with the glands alternating with the sepals in Conceveiba 

 and a few other genera. 2. Pycnocoma contains four well-cha- 

 racterized tropical African species, to which Baillon has added 

 four from Madagascar unknown to me, but for which Mueller has 

 proposed a distinct section, Wetriaria. 3. Ramelia, a New-Cale- 

 donian shrub unknown to me except from Baillon's character, 

 which seems to indicate its place in the present group. 4. Astro- 

 cocca, two species, and 5. Angostyles, one species, all from North 

 Brazil and well defined. 6. Sphcerostyhs, a Madagascar climber, 

 which, from Baillon's character, must differ but little from Flu- 

 tcenetia, except in the calycine segments and stamens reduced to 

 three. 7. PhiJkenetia, about 12 species, dispersed over the tro- 

 pical regions of America, Africa, and Asia, but not extending to 

 Australia. They are mostly climbers, characterized chiefly by 

 the stylar column always fleshy, but varying from globular to long 

 and narrow. Mueller's sections appear to require some modifi- 

 cation. Euplukenetia must be limited to the original P. volubilis, 

 which, besides minor characters, differs very prominently from all 

 others by the very long narrow stylar column crowned by four 

 radiating lobes, although Mueller characterizes the section in 

 which he places it by the stylar column " obovoidea crassa." To 



JjUSTS. JOURN. — BOTAX1T. VOL. XVII. T 



