482 DR. O. STAPF ON SARARANGA SINUOSA. 
Zeit. xxxvi. (1878) t. 10, f. 13). The nucleus was re-absorbed 
almost completely by the embryo sac, even in the youngest 
states which I have seen. In figs. 8 and 9 of Mr. Hemsley’s plate, 
illustrating Sararanga, I have represented this ovary cell as 
communicating with the sutural pores by means of a channel. 
When examining the fresh material from New Georgia I 
was convinced that such a communication does not exist. 
Dr. Guppy’s dried specimens were very much squeezed by 
pressing, hence it was necessary to treat the dissections with 
hydrate of potassium and to flatten them out with needles. In 
doing this I evidently tore the tissue along a line which I, 
biassed by the diagram of the carpel of Pandanus pygmeeus in 
Solms-Lanbach’s paper on Pandanew [Bot. Zeit. (1878) t. 10, 
fig. 25], took to be the stigmatic channel. Thus the gynae- 
ceum appears to be formed by a very great number of carpels 
(Pl. V. figs. 10, 11) which are so completely fused into one 
solid fleshy body as to leave only the stigmas free, and perhaps 
a very small portion below them, where traces of the ventral 
suture, in the shape of an obscure groove, and all but micro- 
scopic pores are visible. 
The anatomical structure of the flowers is on the whole 
very simple, except one or two peculiarities; but as Sararanga 
differs considerably from the rest of the Pandanacew in the 
morphology of its flowers and fruits, it may be worth while 
to mention the principal anatomical features. The epidermis 
of the perianth, as well as of the gynaeceum, consists of poly- 
gonal, slightly oblong, or almost isodiametric cells; the outer 
walls of which are distinctly thicker than the others, and covered 
with a strong and wrinkled or striated cuticle (PI. V., fig. 15; 
VIL, fig. 25). There are stomata present in the perianth (PI. V., 
fig. 15) but none, so far as I am aware, in the gynaeceum. 
The stomata are of the ordinary kind, and surrounded by four 
guard-cells. The succulent parenchyma of the perianth is 
quite homogeneons, except some very large cells which reach 
a maximum length of } mm. and contain bundles of rhaphides. 
The parenchymatic tissue of the gynecium is much more 
differentiated ; but before describing it I will deal with the 
vascular system. A considerable number of vascular bundles 
enter the flower from the pedicel, then, after having given off 
short branches for the perianth, which generally remain simple, 
they divide at the base of the gynaeceum in such a way that 
a 
