DISCHIDIA WITH DOUBLE PITCHERS. 387 
food-substance. From its production in such large quantities in 
the inner pitcher, we may perhaps suppose that this structure, in 
D. pectenoides at least, serves also the purpose of a feeding- 
ground. Whether the fact that the wall of the inner pitcher is 
usually bitten through in one or more places has any connection 
with the presence of this sweet substance, it is impossible to say. 
In this connection attention may be directed to the profuse 
supply of short stalked glands. They appear to be very similar 
in structure to those present in D. Rafflesiana *. With regard to 
the latter the authors state: ‘“ It is quite evident that they have 
nothing to do with any function of the mature pitcher—first, 
because they occur indiscriminately on ordinary leaves and on 
pitchers; and secondly, because they become functionless and 
are cut off by periderm, long before the pitcher is mature” f. 
This may be equally true for the double pitchers. I was, how- 
ever, unable to see that the glands in these pitchers were cut off 
by periderm ; and, as far as I was able to judge from the nature 
of the materials, their appearance suggested no loss of function. 
When the living plant can be studied, the possibility of the 
secretion of these glands being palatable to the ant-inhabitants 
will be worthy of consideration f. 
The mesophyll of the walls of both pitchers consists normally 
of parenchymatous cells whose walls give the cellulose reaction 
with Schulze’s solution. The epidermis (inner and outer) is 
composed of small cells with cuticularized walls, the outer walls 
being for the most part strongly arched and finely grooved. 
Within the mesophyll the much-branched “cells” of the lati- 
ciferous system are abundant (fig. 18, Za.). Microscopic exami- 
nation of the inner surface of the outer pitcher revealed the 
presence of a dense waft of superficial mycelium which was 
easily removed on the point of a needle (fig. 18, myg-). The 
growth of this mycelium appeared to be radial, starting from 
the centre of a curious rosette-like structure (fig. 18, 70.), formed 
by shorter hyphe of a peculiar character. These bore a profuse 
crop of minute abstricted gemme. At the centre of each rosette 
the tissue of the pitcher-wal] appeared to have been punctured. 
A precisely similar mycelium was present on the inner surface of 
* Scott & Sargant, Ann. Bot. vol. vii. pl. 11. fig. 44. 
t Scott & Sargant, /.c. p. 258. ; " 
t “ Most of the Dolichoderi.... lick up the secretions of plants.’ Forel, 
Smithson. Rep. 1894, p. 496. 
