FERTILIZATION OF CLAYTONIA AND PHACELIA. 53 
Tf the flower is fertilized, the stalk bends down through 180° 
until the fruit is ripe, when it once again becomes erect, to 
facilitate the ejection of the seeds*. 
The plants examined bore during the season a number of 
abnormal flowers,with 1, 2, 3, 4, or even all of the stamens aborted, 
and with smaller corollas. This is similar to the phenomenon 
observed in the gynodicecious Labiate +, and it seems therefore 
possible that Claytonia may be gynodiccious, or tending to be- 
come so. Experiments are being made upon this subject. 
PHACELTA. 
Five species of this well-known North-American genus were 
examined, belonging to three subgenera. Gray ¢ arranges the 
genus as follows :— 
§ 1. Evewacetra.—P. tanacetifolia, Benth. 
. CosMANTHUS. 
. COSMANTHOIDES. 
. GYMNOBYTHUS. 
. Wuirtavia, Gray.—P. Whitlavia, Gray, P. Campa- 
nularia, P. Parryi, Torr. 
6. Evtroca.—P. divaricata, Gray. 
7. MicroGENeTES. 
The species examined belong therefore to three out of the 
seven subdivisions of the genus. It is hoped at some future 
time to study examples of the others. As thespecies to be 
here considered differ among themselves, they will be treated 
separately, 
P. tanacetifolia.—The flowers § are borne in dense multiple 
cymes (three or four together), which unroll as they open, keep- 
ing the flowers facing upwards, just as occurs in the Boraginee. 
The flowers are thus massed together, about 10 to 15 open 
flowers occurring at the top of each bundle of cymes. Their colour 
is a pale violet-heliotrope, and there is no perceptible scent, the 
Cw O by 
* Willis, “On the Distribution of the Seed in Claytonia,” Annals of Bot. 
vi. 1892, p. 385. 
t Willis, « Gynodicecism in the Labiate ” (preliminary papers), Proc. Camb. 
Phil. Soe. vii. 1892, p. 348, and viii. 1892, p. 17. 
t ‘Synoptical Flora of North America,’ vol, ii. pt. i- p. 158. 
§ Bot. Reg. t. 1696; Bot. Mag. t. 3703. 
