FERTILIZATION OF PHACELIA. 59 
shape, and versatile, facing upwards. Dehiscence begins near one 
end of each lobe, and the crack spreads rapidly along the entire 
length. Each lobe now turns itself gradually inside out, as one 
might turn a glove-finger, or as occurs in the explosive aril of 
Oxalis, until finally the whole anther looks like a ball of pollen, 
every grain being thus at one stroke extruded from the polien- 
sacs. This is seen in figs. 6 and 7. A similar process occurs in 
Gagea lutea, as described by Kerner *. 
P. divaricata—This species resembles P. tanacetifolia more 
closely than the last three described. The inflorescence is a dense 
cyme, with small flowers. The plant is covered with long silky 
hairs, but is not glandular. The corolla, whose size remains con- 
stant, is about 10 mm. deep and 10 mm. wide, of a funnel-shape, 
and pale heliotrope in colour. The veins are somewhat more deeply 
tinged, and may possibly to some extent act as honey-guides. 
There are longitudinal furrows in the corolla opposite to the 
stamens, but these are not differently coloured from the rest. 
The stamens and style project only slightly beyond the corolla; 
both are hairy, the latter especially. Honey is secreted in the 
usual place, but the disk, instead of secreting it all over its 
surface, bears five well-marked swollen yellow nectaries, alternate 
with the stamens. These latter bear the usual appendages, which 
in this species are adnate also to the corolla, forming little pockets 
at the base of each stamen, between it and the corolla, They 
fit so closely together that there is no room between them for an 
insect to reach the honey, unless it forcibly separate them with 
its proboscis. The style is bifid as usual, but only to a depth 
of about 2 mm. (fig. 18). 
The mechanism of the flower is much like that of the preced- 
ing species. The style, when the anthers open, is about 3-4 mm. 
below, and its lobes are closed against one another. It gradually 
comes up to the same level, and the lobes spread out at an angle 
of trom 60° to 90° with one another. The position of the parts 
favours self-fertilization, while the dichogamy gives the chance 
of across. Insects crawl over the whole mass of flowers as in 
P. tanacetifolia, and probably very often effect autogamy. Every 
flower as a rule sets seed. ; 
Summary: Phacelia.—The above described species present 
similar general features, but differ in detail. Two (P. tanaceti- 
* ‘Pflanzenleben, ii. p. 91. 
