THEIR FORMS AND FUNCTIONS. 487 
pletely free from each other and the petiole. It is worthy of 
mention that in the climbing species of different sections they 
are almost always tolerably broad, in the remainder generally 
linear-subulate.”” Intrastipular thorns also occur, arising from 
intrastipular trichomes. Such are found also in Olneya. 
Bower (on Humboldtia laurifolia, Vahl, in Proc. Phil. Soc. 
Glasgow, xviii. p. 324) describes the peculiar stipules in this 
myrmecophilous plant. They consist of two parts—a lower 
sagittate part with 4-6 glands on its upper surface, and an ovate- 
lanceolate part with one or more glands on the lower surface; they 
originate from only two outgrowths at the base of the leaf-stalk 
before the pinne. 
RosacE&. 
In my previous paper I have already alluded to various inter- 
esting cases presented by species of this order. To these I have 
now several additions. 
In Prunus prostrata, Labill., the stipules are slightly connate, 
converging, intrapetiolar, fringed. Stipules inserted on the 
pedestal of the petiole only, then coming in front of the latter 
they unite with one another, and are therefore practically intra- 
petiolar ; they are connate for a little way only, and then run out 
into two slender, erect, acute, linear, and more or less glandular 
filaments 4-7 mm. long. In the rounded sinus of the fork the 
glands are more or less developed into slender teeth; on the 
lateral and free side of each they are more or less pronounced, 
and one of them (seldom more) runs out into a slender fila- 
ment 1-3 mm. long. This filament recalls the lateral and basal 
spur or branch of the stipules in Lathyrus, and either extends at 
right angles to the main body of the stipules or is clasped round 
the stem. 
The object of these peculiar, intrapetiolar, and connate stipules 
is evidently for the protection of the axillary buds which they 
closely invest. They are also closely lined with silky hairs on 
the inner face; and the lateral filaments or spurs, when rolled 
round the shoot, seem designed to assist in holding the stipules 
in position. The stipules as a whole are more persistent than 
the leaves to which they belong; and as the petiole becomes dis- 
articulated at the point where the stipules cease to be adnate to 
the pedestals, the leaf appears at first sight to be an outgrowth 
