Neopycnocoma. | CXXII. EUPHORBIACE (PRAIN). 963 
ovule. It is, therefore, a matter for further investigation as to whether the 
conditions present do not indicate an abnormal condition of A. macrophylla, Pax, 
showing pistillody, rather than the existence of a distinct genus. 
75a, KLAINEANTHUS, Pierre ex Prain in Kew Bulletin, 1912, 105. 
Flowers dicecious, apetalous. Male: Sepals 4—5, imbricate, connate 
below in a campanulate tube as long as the lobes. Stamens 8 or 10, 
2-seriate ; filaments of the outer series rather shorter than those of the 
inner, surrounded by a ring of 4-5 fleshy ovate glands opposite the 
sepals and alternate with the outer stamens; anthers basifixed, smooth, 
2-celled, dehiscence longitudinal, introrse. Rudimentary ovary 2-3-lobed, 
as long as the stamens of the inner series, Female: Sepals 5, rarely 4, 
imbricate, almost free to the base. Disk cupular with often an incom- 
plete or complete ring of 5 or 4 staminodes inserted at the base of the 
disk within. Ovary 3-celled, glabrous; styles 3, glabrous, 2-partite to 
their lower fourth, each arm obtusely 2-lobed at the tip. Fruit capsular, 
3-coccous ; cocci crepitant, separating septicidally and also opening loculi- 
cidally leaving a long trigonous persistent woody columella; epicarp 
thinly fleshy ; endocarp thickly woody. Seeds large, flattened ovoid, 
carunculate; embryo rather large; cotyledons broad, ovate-cordate, 
flat A tree; leaves alternate; petioles pulvinate at the base and 
apex; flowers in ample terminal and axillary panicles; males glomeru- 
late, females subsolitary to a bract ; pedicels in both sexes slender, short, 
articulate. 
A single endemic species. 
1. K. Gabonize, Pierre ex Prain in Kew Bulletin, 1912, 106. A 
tree 30-60 ft. high, twigs reddish and pubescent when young, at length 
grey and glabrous. Leaves distinctly petioled, oblong or obovate, bluntly 
acuminate, margin entire, base wide- to narrow-cuneate, 4-8 in. long, 
13-3} in. wide, firm, dark green and dull above, paler and somewhat 
polished beneath, glabrous on both surfaces ; secondary nerves 8-12 
pairs, raised beneath, looping within the margin ; petiole slightly pulvi- 
nate at the base, distinctly pulvinate and geniculate at the apex, 1}-2 in. 
long. Panicles axillary and terminal, males as long as the females, 
rather shorter ‘than the leaves; male flowers glomerate, females sub- 
Solitary ; pedicels in both sexes shorter than the calyx, distinctly articu- 
late. Male: Calyx 5- or 4-lobed, companulate below; lobes oblong, 
obtuse, pubescent and ciliate, strongly imbricate. Stamens 8 or 10, 
2-seriate ; filaments of outer series shortest, surrounded by a ring of 4 
or 5 extra-staminal glands alternate with the outer stamens and opposite 
the calyx-lobes ; anthers basifixed, erect, introrse, dehiscence longitudinal. 
Rudimentary ovary 3-gonous, as long as the inner stamens, = (rarely 2-) 
lobed at the apex. Female: Calyx 5- (less often 4-) partite almost to 
the base, otherwise as in the males. Disk cupular, denticulate and 4-5- 
lobed. Staminodes often present in a complete or incomplete series, 
hypogynous. Ovary 3-celled, glabrous; styles 3, deeply 2-partite, the 
arms again minutely and bluntly 2-fid. Capsule 3-celled, } in. across ; 
