320 
Angelica, Geyer having previously sent it to him as a species of Thas- 
pium. Walper! follows Hooker in retaining the species in Angelica. 
Mr. Watson? first calls attention to the improbability of its being an 
Angelica. No additional information could be obtained when Coul- 
ter and Rose’s Revision of Umbelliferze was written, and so the species 
was again doubtfully referred to Angelica. Geyer collected his plant in 
the Nez Perces Indian Reservation in 1841 (?), and, since no botanical 
collector had visited this region later, it was impossible to associate 
the description with any known species. The habit of the plant sug- 
gested to me Ligusticum, but I repeatedly failed to convince myself 
that it belonged in that genus. In 1892 Messrs. Sandberg, MacDougal, 
and Heller spent some time on the Nez Perces Indian Reservation, and 
it was therefore with considerable interest that I looked over their col- 
lection of Umbellifere. It was found to contain a plant which is un- 
doubtedly the Angelica verticillata of Hooker and which must be a 
Ligusticum. We are accordingly warranted in adopting the following: 
Ligusticum verticillatum Hook.) Coult. & Rose; Angelica verticillata Hook. Lond. 
Journ. Bot. vi, 233 (1847). PLATE XII. 
Hooker’s notes and description are as follows: 
‘Shady grassy borders of pine woods, on high plains of the Nez Perces Indians. 
June (n. 414), Of the genus of this lam exceedingly doubtful. The immature fruit 
and leaves are not unlike those of some Angelica; but there are no large sheathing 
bases to the petioles. My specimens are very imperfect. They consist of a fusiform 
root, clothed, especially above, with dense, coarse, long fibers, the remains of former 
petioles; there is only one root leaf, the main petiole of which is about a span high, 
semiterete, channeled, striated, glabrous, as is the whole plant. This divides into 
3 at the top, and each of those again inte 3, bearing generally 5 oblong-ovate, 
membranous leaflets, opposite and slightly petiolate; the extremity of a flowering 
branch has a whorl of about 4 pinnated and laciniated, sessile, small (2 to 3 inches 
long) leaves, within which is an umbel of 6 rays; the centerray is twice as long and 
4 or 5 times as stout as the others, each bearing an umbel of many petiolated umbel- 
lules of fertile flowers, yellowish-white, the stylopodia very large, much broader 
than the ovary; the other 5 rays are sterile (having no trace of pistil), and are each 
terminated by a compound sterile umbel, the whole forming a whorl around the cen- 
tral ray, whence Mr. Geyer’s specific name. There is besides a separate fertile 
umbel, with very immature, deeply sulcated, obovate fruit. The umbels and umbel- 
lules have no involucral scales. ” 
EXPLANATION OF PLATE.—Fig. a, dorsal view of carpel, enlarged 5 ciameters; b, cross section of 
carpel, enlarged 10 diameters. 
Ligusticum eastwoode Coult. & Rose, sp. nov. Piatr XIII. 
Low acaulescent plant from a short caudex; scape 10 to 30 cm, high, glabrous; 
leaves pinnate, 10 to 15 cm. long; pinniw 7 to 13, opposite, oval, 2- to 3-lobed and 
lobes 2-to 3-cleft into linear acute segments; inflorescence very much congested, 
almost head-like; involucre 1- to 2-leafed or none; bractlets of the involucels con- 
spicuous, longer than the fruit, 2- to 5-toothed; rays few, 2 to 3 lines long; pedicels 
about 1 line long; fruit ovate, glabrous, flattened laterally, 3 mm. long; carpels 
flattened dorsally; ribs 5, small; oil-tubes 2 to 3 in the intervals; seed face with a 
broad, shallow concavity; calyx teeth prominent; stylopodium conical. 
‘Annales, i, 350. 2?Proc. Amer. Acad, xvii, 374. 
