PITTIER TREATMENT OF THE GENUS CASTILLA. 



273 



Receptacles of the primary stamiuate inflorescence stipitate, flabellate, in 

 axillary clusters generally of 0. geminate, the two exterior pairs flowering 

 first. Basal bracts small (3 to 4 mm. long), ovate-acuminate, hairy outside, 

 smooth inside, involucrate. Stipes about 2 cm, long, hairy and bearing a few 

 more or less developed bracts. Lobes of receptacle 2 to 3 cm, long, 2 cm. broad, 

 kidney-shaped, flat cr reflexed. Scales in 5 or 6 imbricate rows, 2 to 3 mm. 

 long and broad, ovate-acuminate, hairy- Interstaminato bracts small (about 

 1 mm. long), ovate-lanceolate or acuminate. Stamens 2 to 4 mm. long, glabrous 

 and numerous; filaments ratber slender; connective thick, papillose on the 

 upper face: anther cells apparently broader than in the other species. 



Receptacles of the complemental staminate inflorescence geminate, stipitate, 

 claviform. 2 to 2.5 cm. long, with a slit-like opening. Stipes 1 to 1.5 cm. long, 

 thick, pubescent, squamose. Scales broadly ovate (2.5 

 to 4 mm. long, 4 to G mm. broad), pubescent, almost 

 free to the base. Interslaminate bracts hairy on both 

 faces and filiate, either simple and narrow, or broad 

 and diversely laciniate. Stamens glabrous. 1 to 5 mm. 

 long. 



Receptacles of the pistillate inflorescence sessile or 

 borne on a very short, thick peduncle, cupuliform, cov- 

 ered outside by about 5 rows of imbricate scales, ovate- 

 acuminate, hairy outside. Perianth about 3 mm. long, 

 free to the base, longitudinally 4 or 5-sulcate, divided 

 at the tip into 4 or 5 uneven, more or less acute lobes. 

 Style 1.5 to 2 nun. long, rather thick, more or less cov- 

 ered with stiff hairs at the base and along one side; 

 stigmas 2. seldom 3, 2 to 4 mm. long, linguiform, 

 papillose, purplish or greenish yellow, semierect and 

 more or less twisted. 6 



Fruiting receptacles very much enlarged and reach- 

 ing often more than 5 cm. in diameter. Achenia 14 to 

 22 on each receptacle, 18 to 20 mm. long, 12 to 15 mm. 

 in diameter, orange-red at maturity, scarcely concres- 

 cent at the base, but simply pressed together without. 



losing their individuality, and always free at the apex. Seed ovoid with more 

 or less flattened sides. 11.3 mm. long, 8.4 nun. in diameter; episperm white, 

 arillus brown. 



Type in the U. S. National Herbarium, nos. 408077-8, collected by O. F. Cook 

 (no. 205), March 25), 1902, at Secanquim, Alta Verapaz, Guatemala, altitude 

 550 meters. 



Fig. 51. — Floral details 

 of Castillo, guatema- 

 lensis. a, Stamen, 

 front view, showing 

 large connective; b, 

 .stamen, back view ; c, 

 group of stamens at 

 various stages, with 

 bracteoles also ; tl, d' , 

 d" , styles. Scale 3. 



"The foliaceous radial crests along which the stamens are usually inserted 

 are here little and irregularly developed, being at times narrow, either smooth 

 or ciliate, and at other times broader and with triangular or rounded, ciliate 

 teeth; bracteoles are never seen high on the filaments; all transitional stages 

 between a bract and a perfect stamen' can be found on one receptacle. The 

 stamens develop successively from the base of the lobe toward its periphery, 

 the basal becoming fully mature and having their anther cells open and empty 

 while the marginal ones are still very short, with the connective inflexed on the 

 filament. 



6 In Secanquim dry materials, the pistil had taken the appearance of a folia- 

 ceous organ, opening by a side slit, and then forming a hilobulate or trilobulate 

 leaflet, with a distinctly marked midrib. In the specimen with two styles the 

 hatry external zone was found to correspond with the broad interlobular rib. 



