SAFFORD—-CLASSIFICATION OF ANNONA. 7 
Leaves without pockets in the axils of the nerves, 
Blades flat, thick, coriaceous, the lateral nerves 
impressed on both surfaces______---_-_-. 2. PSAMMOGENIA (p. 8). 
Blades more or less conduplicate and undulate, 
submembranaceous, the lateral nerves 
impressed above, very prominent beneath. 3. ULocarrus (p. 9). 
Section 1. EUANNONA Safford. 
(Guanabani Mart. in part. Euannona Safford, Journ. Washington Acad. Sci. 
1: 118. 1911, restricted.) 
Type species, Annona muricata L. (pl. 1). 
This section is distinguished from all other sections of the genus Annona by 
minute pits or pockets in the axils of the lateral nerves of the leaves (fig. 5), 
sometimes quite conspicuous, as in the 
case of Annona montana Macfad., but 
often scarcely visible to the naked 
eye. The peduncles are solitary or 
geminate, in the type species fre- 
quently caulifloral, the flowers large, 
6-petaled, the 3 outer petals thick, 
valvate, broadly ovate, usually cor- 
date at the base and acute or acumi- 
nate at the apex. The 3 inner petals 
are somewhat smaller and thinner, 
concave, with their edges imbricate 
or overlapping and forming a dome- 
like covering for the essential parts 
(pl. 6), obovate or suborbicular, ob- 
tuse or rounded at the apex, and 
Fig. 5.—Leaf of Annona mon- 
tana, showing axillary Fic. 6.—Annona montana. Flower, fruit, and 
pockets. Scale 4. leaves. Scale 4. 
usually clawed at the base. ‘The ovaries are linear and clothed with hairs, a 
solitary ovule at the base and an ovoid or oblong style at the apex (fig. 3, b), 
and are quite distinct (discrete) in the flower (pl. 3). The fleshy fruit (syn- 
carpium) is muricate with fleshy prickles (fig. 6), consisting of the persistent 
tips of the ovaries from which the jointed styles become detached soon after 
pollination. 
In addition to Annona muricata L., which has been cultivated from pre- 
historic times for the sake of its pleasantly acidulous, juicy fruit, this section 
includes several very closely related wild species, the first of which to receive 
a binary Latin name in accordance with botanical usage was Annona montana 
EXPLANATION OF PLATE 3.—Showing discrete ovaries of the section Euannona. Photo- 
graphed from material in the U. S. National Herbarium by W.S. Clime, Scale 6. 
11419°—14——2 
