STANDLEY—TROPICAL AMERICAN PHANEROGAMS. 125 
Duggena Vahl, published in 1793, should not be applied to the group. 
Its type is Duggena richardii Vahl, which apparently is a synonym of 
D. spicata, listed below. | 
Four species of the genus occur in Panama: D. ovatifolia, described 
from Costa Rica; D. panamensis, first characterized by Cavanilles 
in 1801 from specimens collected on Ancén Hill, Panama; D. hayesi, 
also described from’ Panama; and a fourth, collected recently by Mr. 
Pittier, which appears to be undescribed. 
Duggena rudis Standley, sp. nov. 
Branches slender, brownish, densely strigose with whitish tawny hairs; stipules 
4 to 6 mm. long, triangular, with long filiform tips; leaves nearly sessile, the petioles 
1 to 2mm. long, the blades ovate or oblong-ovate, 35 to 55 mm. long, 14 to 24 mm. wide, 
rather abruptly acuminate, rounded at the base, thin, bright green, strigose on the 
upper surface, more densely pubescent beneath, especially along the veins, with 
appressed whitish hairs; racemes slender, 6 to 9 cm. long; flowers about 3 in each 
cluster, on very short peduncles; pedicels about 1 mm. long; bracts linear-subulate, 
3 mm. long or less; corolla not seen; fruit 4-celled, depressed-globose, 2.5 mm. in diam- 
eter, densely hirsute; calyx lobes persistent, lance-linear, attenuate, nearly as long 
as the fruit, sparingly pubescent outside, glabrous within. 
Type in the U. 8. National Herbarium, no. 676602, collected in shady places along 
the Chagres River, near El Vigfa, Panama, January 12, 1911, by H. Pittier (no, 237 8). 
Resembling Duggena panamensis in the 4-celled fruit, but readily distinguished by 
the small, nearly sessile leaves, narrow and longer calyx lobes, and densely pubes- 
cent fruit. 
Duggena asperula (Wernham) Standley. 
Gonzalea asperula Wernham, Journ. Bot, Brit. & For. 51: 219. 1913, 
Duggena brachyantha (A. Rich.) Standley. 
Gonzalea brachyantha A. Rich. in Sagra, Hist. Cuba 11: 16. 1850. 
Gonzalagunia brachyantha Urban, Symb. Antill. 7: 400. 1912. 
Duggena bracteosa (Donn. Smith) Standley. 
Gonzalea bracteosa Donn. Smith, Bot. Gaz. 33: 252. 1902. 
Gonzalagunia bracteosa Robinson, Proc. Amer. Acad. 45: 405, 1910. 
Duggena grisea (Wernham) Standley. 
Gonzalea grisea Wernham, Journ. Bot. Brit. & For. 51: 218. 19138. 
Duggena hayesii (Wernham) Standley. 
Gonzalea hayesii Wernham, Journ. Bot. Brit. & For. 51: 219, 1913. 
Duggena leptantha (A. Rich.) Standley. 
Gonzalea leptantha A, Rich. in Sagra, Hist. Cuba 11: 16. 1850. 
Gonzalagunia leptantha Urban, Symb. Antill. 7: 400. 1912. 
Duggena mollis (Spruce) Standley. 
Gonzalagunia mollis Spruce; Schum, in Mart. Fl. Bras. 6°: 290. 1889. 
Gonzalea mollis Spruce; Schum. in Mart. Fl. Bras. 6°: 291. 1889, as synonym ; 
Wernham, Journ. Bot. Brit. & For. 51: 219. 1913. 
The type collection is the same in both places of publication cited above; Mr. 
Wernham, however, makes no reference to the earlier publication of the species by 
Schumann. 
Duggena nivea (Bartling) Standley. 
Gonzalea nivea Bartling; DC. Prodr. 4: 436. 1830. 
Gonzalagunia nivea Kuntze, Rev. Gen. Pl. 1: 284. 1891. 
