236 CONTRIBUTIONS FROM THE NATIONAL HERBARIUM. 
Coumarouna panamensis Pittier, sp. nov. 
Section Eucoumarouna: A portly tree, often reaching 50 meters in height 
and 1 meter in diameter at the base. Crown rounded. Bark brownish, slightly 
scaly. Twigs densely lenticellate. 
Leaves alternate, glabrous, 5 to 8-foliolate. Petiole 40 to 70 cm. long, flat- 
tened above, 6 to 10 mm. broad at the base, closely reflexed-winged on both 
sides through its whole length, with a median ridge on the upper face from 
the first leaflets to the end; lower, rounded side densely lJenticellate. First 
pair of leaflets opposite, the others alternate and more spaced along the middle 
part of the main rachis. Leaflets petiolulate, strongly inequilateral or dimidiate, 
discolored, densely covered with transparent dots; petiolules 10 to 15 mm. 
long, sharp-edged on the lower side; leaflet blades elliptic, oblique, 15 to 80 
cm. long, 5 to 7 cm. broad (the narrow side equal to about one-third of the 
whole breadth), subacute at the base, subobtuse at the tip; nervation salient 
beneath with the costa acutely carinate. Stipules very caducous; stipels 
lanceolate, 4 to 7 mm. long. 
Panicles terminal, loosely branching, 80 to 40 cm. long. Rachis more or 
less pubescent. Flowers large, pink, pedicellate. Pedicels 4 to 5 mm. long, 
minutely pubescent. Bracts and bractlets very small, hairy, deciduous. Calyx 
tubular, attenuate at the base, pubescent, the tube about 6 mm. long, the two 
anterior lobules 16 to 17 mm. long, 7 mm. broad, glandular-dotted, the three 
posterior acute and small (the middle one larger, about 1 mm. long). Vexillum 
suborbicular or ovate, hardly as long as the anterior calyx lobes, bilobate at 
tip. Wings obovate-oblique, about 16 mm. long, 6.5 mm. broad; the inferior 
lobe shorter and overlapping the superior one; claw slender, 1.5 mm. long. 
Petals of the keel free, about 19 mm. long and 6.5 mm. broad, minutely ciliate 
on the carinal margin, auriculate at the base. Stamens almost equal, all 
connate nearly to the tip; anthers ovate, short-apiculate. Ovary glabrous, 
uniovulate. 
Fruit flat, elliptic, 6 cm. long, 3.5 cm. broad, and 8 em, thick or less, 
pedicellate, rounded and obscurely apiculate at tip, hairy outside in the fresh 
condition, chocolate-brown and glabrous when dry, the transverse section of 
the interior cavity subcircular. Pedicel about 1.5 cm. long, thick. Seed 
almond-like, straight, elongate-oblong, and compressed, 5 cm. long, 1.8 em. 
broad, with a light brown episperm. 
Type in the U. S. National Herbarium, no. 679031, collected at Bailamonos, 
between Mamei and San Pablo, Canal Zone, Panama, on a hillside, flowers, 
July 12, 1911, by EB. D. Christopherson (Pittier, no. 8950). 
OTHER COLLECTIONS: 
PANAMA: Along the Trinidad River, Canal Zone, near sea level, flowers, 
July 19, 1911, Pittier 4033. Forests of Pauarandé6, Sambfi Valley, 
southern Darién, fruits, February 8, 1912, Pittier 5727. 
Coumarouna panamensis is nearly related to C. oleifera (Benth.) Taub., of 
the Mosquito Coast, a species imperfectly described. It has the same large, 
glandular calycinal wings, but differs in the shape of the vexillum and wings, 
in the longer and more rounded fruits, and in the size and shape of the seeds. 
The leaves of the latter species have not been described. 
The fresh fruits of C. panamensis are covered with a grayish green pubescence 
and impregnated with an oily, sweet-smelling fluid, which may have its origin 
in the resinous channels that crowd the mesocarp. When the fruits are dry, 
these ducts are seen filled with a crystalline substance. 
