an eel 
REVISION OF OYEDABA. 417 
BLAKE 
SPECIMENS EXAMINED: 
Bortvia: Apolo, altitude 1,465 meters, 1902, Williams 317 (B, N, photo. and 
fragm. G); July 9, 1902, Williams 1408 (type; N. Y. Bot. Gard.). 
This species is distinguished from 0. rusbyt, its nearest relative, by its more lanceo- 
late leaves, which are less densely pubescent beneath. 
7. Oyedaea boliviana Britton, Bull. Torrey Club 19: 149. 1892. 
Helianthus mandonii Schultz Bip. Bull. Soc. Bot. France 12: 79.1865, nomen nudum. 
Oyedaea pearcei Rusby, Mem. Torrey Club 3: 59. 1893. 
Evergreen shrub, 2 to 2.6 meters high, much branched; stem stout, terete, fuscous- 
brown, rather densely hispidulous or hispid-pilose with spreading or incurved hairs 
and sordid-glandular, especially on the younger parts, in age subglabrate but 
tuberculate below; leaf blades 8.5 to 17 cm. long, 1.3 to 7.5 cm. wide, ovate or 
broadly ovate to lance-ovate or the uppermost lanceolate, acuminate, cuneate to 
rounded-cuneate at base, triplinerved (sometimes obscurely 80) and rather veiny, 
obscurely appressed-serrulate (the teeth mucronate, 29 pairs or less) or the upper 
subentire, above dark green, hispidulous and hispid with incurved hairs with 
tuberculate persistent bases, in age subglabrate, beneath slightly paler, rather 
densely hispidulous-pilosulous with incurved-spreading, harsh or soft hairs with 
scarcely swollen bases, more or less hispid along the veins, and gland-dotted; upper 
leaves much reduced; petioles 4 to 18 mm. long, pubescent like the stem and 
distinctly ciliate; heads rather numerous, 1.8 to 3 em. wide, ternately arranged at 
ends of branches and branchlets, on pedicels 2 to 28 mm. long or less; disk 7.5 to 
10 mm. high, 7 to 10 mm. thick; involucre 3 to 4-seriate, graduated, 4.5 to 7 mm. 
high; phyllaries ovate to oblong-ovate, with pale, strongly indurate and thickened, 
mostly 1-vittate base, more or less hispidulous and hispid with appressed hairs chiefly 
along middle and on margin (where the hairs are spreading) or sometimes all over, 
and shorter, acute or acuminate, deltoid or triangular, loose or reflexed, herbaceous 
apex, the innermost with drier paler tips; receptacle flattish; rays about 12, oval to 
linear-oblong, hispidulous on nerves of back, 8 to 18 mm. long, 2.5 to 4 mm. wide; 
disk corollas slender, puberulous on teeth, otherwise glabrous, 4.5 to 6.8 mm. long 
(tube 1.5 to 2.2 mm.); pales narrow, carinate, puberulous and gland-dotted at apex, 
acutish to acuminate, 7 to 8 mm. long; achenes (nearly mature) flattish, blackish 
brown, sparsely strigose on sides, ciliate on the narrow wings, 3.5 mm. long, con- 
tracted above into a more broadly winged short neck; awns 2, unequal, 3.7 mm. 
long or less, decurrent into the wings; corona 0.7 mm. high, of about 4 pairs of 
squamellae united to above the middle. 
Type LOCALITY: Yungas, Bolivia. 
SPECIMENS EXAMINED: 
Boutv1a: Yungas, altitude 1,830 meters, 1885, Rusby 2143 (type of O. boliviana; 
herb. Columb. Coll.); in 1890, Bang 546 (type collection of O. pearcet; 
B, G, K, N). Santa Cruz, July, 1865, Pearce (B, K, fragm. G.). Vicinity 
of Yungas, altitude 2,100 meters, 1906, Buchtien 190 (N. Y. Bot. Gard.). 
Polo-polo near Coroico, altitude 1,100 meters, 1912, Buchtien 3945 (N. Y. 
Bot. Gard.). Near Ananea, Province of Larecaja, January, 1860, Mandon 37 
(type collection of Helianthus mandonit; B, fragm. G). Near Sorata, 
Province of Larecaja, Mandon (G). 
This species is usually readily distinguishable from its relatives by its distinctly 
triplinerved leaves. Although it is somewhat variable in leaf shape, it is impossible 
to recognize more than one species in the moderately extensive material I have 
examined. The reference of this species to Helianthus by Schultz Bipontinus, in 
his enumeration of Mandon’s Bolivian Compositae, is an indication of the haste or 
carelessness with which that usually well-informed specialist must have made his 
determinations of those plants. 
