BLAKE—REVISION OF TITHONIA. 433 
Oaxaca: Vicinity of Totontepec, altitude 1,675 to 2,135 meters, 1894, Nelson 
772 (N). 
Curapas: Between San Sebastién and Pantepec, 1907, Collins & Doyle 211 (N). 
Guatemata: Cob4n, altitude 1,350 meters, 1908, Tiirckheim II. 2053 (N). Near 
Santa Marfa, Department of Quezaltenango, altitude 1,500 to 1,800 meters, 
1905, Maron & Hay 3595 (N). Quezaltenango, altitude 1,980 meters, 1917, 
Holway 729 (G). 
Satvapor: Opaneca, Department of Ohuachapan, altitude 1,460 meters, 1907, 
Pittier 2007 (N). 
Honpuras: Between Llano de la Puerta and El Salto, Copdn, altitude 900 
meters, 1907, Pittier 1854 (N). 
Costa Rica: Along the Rfo Tiliri, 1892, Tonduz 7196 (N). 
Tonduz 7196 is peculiar in having the leaves beneath green, merely hispidulous- 
puberulous, and rather densely gland-dotted, in this respect approaching T. calva 
lancifolia, but it is otherwise typical of T. scaberrima and possesses the characteristic 
involucre of the latter species. T'. scaberrima is readily told by its densely pubescent 
stem and leaves, large heads, and large, broadly rounded phyllaries. 
9. Tithonia fruticosa Canby & Rose, Contr. U. S. Nat. Herb. 1: 104. pl. 5. 1891 
Stout shrub, 3 to 4 meters high, the woody stem becoming 10 cm. in diameter, 
glabrate and clothed with a grayish bark; younger branches densely and canescently 
pilose-tomentose with spreading hairs; leaves opposite below, alternate above, the 
blades 6.5 to 30 cm. long, 2.2 to 14 cm. wide, ovate or lanceolate, acuminate or attenuate 
and often falcate, cuneate into the margined petiole, crenate-serrate, triplinerved and 
veiny beneath, canescent-pilose on both sides but particularly beneath with rather soft 
incurved tuberculate-based hairs and gland-dotted; petioles 3 to 5 cm. long, margined 
but not auriculate; heads terminal and axillary, 7 to 9.5 cm. wide, on fistulose striate 
peduncles shorter than the leaves; disk 2 to2.5 cm. high, 2.5 to4.5 cm. thick; involucre 
4-seriate, strongly graduated, 2 to2.8 cm. high, the phyllaries broadly oval or obovate- 
oval (the median 8 to 12 mm. wide), broadly round-tipped or rarely obtuse, the outer 
densely canescent-pilosulous with tuberculate-based appressed hairs, the inmost less 
pubescent and greenish, all with obscurely indurate base and shorter submembrana- 
ceous tip, the inmost with submembranaceous-subherbaceous tip; rays about 14 to 20, 
yellow, narrowly elliptic, 2 to 3.5 cm. long; disk corollas hispidulous throughout, or 
only below and on the teeth, 8 to 9 mm. long (tube 1 mm.); pales ovate-lanceolate, 
stiff, abruptly pointed but not cuspidate, finely tuberculate-hispidulous and gland- 
dotted above or subglabrate, 12 to 14 mm. long; achenes appressed-pilose, 4 to 5 mm. 
long; squamellae connate into a laciniate-fimbriate paleaceous crown 1 to 1.5 mm. high, 
the awns obsolete or represented merely by teeth only twice as long as the crown. 
Type LocaLiry: Alamos, Sonora. 
ILLUSTRATION: Contr. U. S. Nat. Herb. 1: pl. 5. 1891. 
SPECIMENS EXAMINED: 
Sonora: Among bushes near a watercourse, Alamos, March or April, 1890, Pal- 
mer 303 (type collection; G. N). Huehuerachi, altitude 1,220 meters, 1890, 
Hartman 301 (G, N). High up on the Sierra de Alamos, 1910, Rose, Standley 
& Russell 13085 (N). 
Curmvuanua: Near Batopilas, 1898, Goldman 232 (N). 
Duranco: San Ramén, 1906, Palmer 68 (G, N). 
Sivatoa: Arroyo de la Labor, San Ignacio, altitude 380 meters, 1918, Montes & 
Salazar 309 (N). 
This very distinct species is quickly recognized by its shrubby habit, densely 
canescent-pubescent stem and leaves, and large rounded phyllaries. 
