506 ESCALLONIACEAE 
1. HYDRANGEA L. 
Shrubs or trees. Leaves opposite, without stipules: blades entire, toothed, or lobed. 
Flowers perfect, except a few conspicuous ones around the edge of the corymb or thyrse. 
Sepals united into a truncate or 4—5-toothed border, or in sterile flowers of 4-5 spreading 
petal-like lobes. Petals 4-5, valvate. Stamens 8-10, inserted at the base of an epigynous 
disk : filaments filiform. Ovary inferior, completely or partially 2-4-celled : styles 2-4, 
distinct or slightly united at the base. Ovules numerous, borne on axile placentae. 
Capsule 2-4-celled, membranous, opening at the top between the styles. Seeds numerous, 
minute. Testa membranous, reticulated. The plants flower during the spring and summer. 
Leaf-blades merely toothed : inflorescence corymbose. 
Leaf-blades glabrous or inconspicuously pubescent beneath. 1. H. arborescens. 
Leaf-blades conspicuously and copiously pubescent beneath. 
Leaf-blades membranous, light gray beneath. 2. H. cinerea. 
Leaf-blades leathery, bright white beneath. 3. H. radiata. 
Leaf-blades deeply lobed : inflorescence thyrsoid. . 4. H. quercifolia. 
1. Hydrangea arboréscens L. A branching, sometimes straggling shrub 1-3 m. tall. 
Leaf-blades ovate, oval or elliptic, 0.5-2 dm. long, short-acuminate, serrate, acute, rounded 
or cordate at the base, glabrous or sometimes minutely pubescent beneath ; petioles shorter 
than the blades: corymbs 5-20 cm. broad, usually with few sterile ray-flowers or these 
wanting, commonly pubescent: hypanthium campanulate, 1-1.5 mm. broad : sepals minute, 
triangular: petals deciduous, concave, ovate, about 1 mm. long: stamens conspicuously 
exserted : stigmas reddish : capsules 2 mm. broad, 8-ribbed, broader than high, each tipped 
with the 2 spreading styles. 
In woods or along streams, New York to Iowa, Florida and Louisiana. 
2. Hydrangea cinérea Small. A spreading shrub, 1-2 m. tall, with deep green 
foliage. Leaf-blades thinnish, oval, elliptic, narrowly ovate or orbicular-ovate, 6-15 cm. 
long, acuminate, serrate, obtuse, rounded or cordate at the base, bright green above, gray- 
tomentose and not reticulated beneath ; petioles somewhat more than one-half as long as 
the blades or shorter: corymbs 5-15 em. broad, rather round-topped : sterile ray-flowers 
commonly present: hypanthium campanulate, ribbed : sepals triangular, acute: petals 5, 
ovate, boat-shaped, 1.5 mm. long, hooded at the apex: stamens conspicuously exserted : 
capsules urn-shaped,. about 2 mm. in diameter, usually higher than broad, strongly ribbed, 
each tipped with the 2-3 spreading styles. 
On the slopes of the Alleghenies, North Carolina, Tennessee, Georgia and Alabama. 
3. Hydrangea radiàta Walt. A branching shrub, 1-2.5 m. tall. Leaf-blades leathery, 
ovate, varying to oval or ovate-lanceolate, 6-12 cm. long, acuminate, serrate or crenate- 
serrate, acute, rounded or subcordate at the base, bright green and glabrous above, densely 
silvery white-tomentose and reticulated beneath ; petioles about 3 as long as the blades or 
shorter: corymb round-topped, 4-12 cm. broad: sterile flowers always present: hypan- 
thium urn-shaped, ribbed: sepals triangular or usually somewhat acuminate: petals 5, 
ovate, concave, nearly 1 mm. long, apiculate: stamens much exserted: capsules urn- 
shaped, 2 mm. broad, strongly ribbed, shorter than their pedicels, each tipped with the 2 
spreading swollen styles. 
On rocky slopes, North and South Carolina. 
4. Hydrangea quercifólia Bartr. A stout shrub, 1-2 m. tall, with densely tomentose 
twigs and branches. Leaf-blades oval or suborbicular in outline, usually 5-lobed, 6-20 
cm. long, tomentose beneath, the lobes serrate and often slightly lobed ; petioles ri M 
long as the blades : panicles 1-2 dm. long : sterile flowers numerous, white, turning purple : 
hypanthium urn-shaped, longer than the pedicel: sepals ovate, about 4 as long as the 
hypanthium: petals 5, oblong, 1.5 mm. long, slightly concave: stamens conspicuously 
exserted : capsules urn-shaped, 2.5 mm. in diameter, ribbed, each tipped with the spreading 
much enlarged styles. 
On river banks, middle Georgia to Florida and Mississippi. GRAY-BEARD. OLD MAN’s-BEARD. 
2. PHILADÉLPHUS L.! 
Shrubs, with branching stems. Leaves opposite, deciduous: blades simple, petioled : 
stipules wanting. Flowers solitary, corymbose or racemose, either terminal or axillary. 
Hypanthium turbinate, adnate with the ovary. Sepals 4 or 5, persistent. Corolla white 
or cream-colored. Petals 4 or 5, obovate or rounded, convolute in the bud. Stamens 
numerous, inserted on the disk : filaments slender. Styles 3-5, united, at least at the base : 
! Contributed by Mr. C. D. Beadle. 
