PRIMULACEAE 901 
many stamens as there are corolla-lobes and opposite them, adnate to the tube 
or on the calyx, rarely alternating with staminodia. Filaments sometimes united 
at the base. Anthers 2-celled: sacs opening introrsely. Gynoecium a single 
pistil. Ovary free or nearly so, 1-celled, with a free central placenta. Style 
entire. Stigma entire or lobed. Ovules amphitropous, numerous or two. 
Fruit a 1-celled capsule opening by 2 to 8 valves. Seeds 2 or many, with a 
smooth roughened or furrowed testa. Endosperm fleshy or horny. Embryo 
usually transverse. 
Corolla-lobes erect or spreading. 
Lobes of the corolla imbricated. 
Ovary superior: plants with dissected leaf-blades. 1. HOTTONIA. 
Ovary partly inferior : with entire leaf-blades. 2. SAMOLUS. 
Lobes of the corolla valvate or convolute. 
Capsules opening lengthwise. 
Staminodia wanting: filaments united at the base. 3. LYSIMACHIA. 
Staminodia present : filaments distinct or essentially 80. 4. STEIRONEMA. 
Capsules circumscissile. 
Corolla longer than the calyx : filaments pubescent, adnate to the baseof the 
corolla. 5. ANAGALLIS. 
Corolla shorter than the calyx: filaments glabrous, adnate to the corolla- 
tube. 6. CENTUNOULUS. 
Corolla-lobes reflexed. . DODECATHEON. 
~J 
l. HOTTÒNIA L. 
Aquatic herbs, with glabrous foliage. Stems submerged. Leaves numerous, the sub- 
mersed with pinnately dissected blades : petioles often inflated. Scapes erect, the inter- 
nodes inflated. Bracts whorled. Flowers whorled at the nodes, pedicelled. Calyx persist- 
ent: lobes 5, narrow, imbricated, longer than the tube. Corolla white, salverform : 
tube longer than the 5 imbricated lobes of the spreading limb. Stamens 5, adnate to the 
corolla-tube below the middle: filaments about as long as the anthers. Ovary sessile: 
style filiform: stigma capitate. Ovules numerous, anatropous, inserted on a rounded 
stalked placenta. Capsule membranous, partially 5-valved. Seeds numerous, slightly 
angular. FEATHERFOIL. 
1. Hottonia inflata Ell. Stems submerged, spongy, 2-6 dm. long, leafy, simple or 
branching : leaf-blades ovate to oblong in outline, 5-10 cm. long, finely divided almost to 
the middle into linear-filiform segments: scapes tufted, 1-3 dm. long, jointed, the lower 
joint often 2.5 cm. thick, the upper ones successively smaller: bracts linear or linear-spat- 
ulate, 5-15 mm. long: flowers whorled at the nodes: pedicels variable in length, 0.3-5 cm. 
long: calyx often minutely glandular; tube very short; lobes linear, 5-10 mm. long, 
rather obtuse: corolla white, 3 mm. long, shorter than the calyx, the lobes oblong, obtuse, 
3 aslong as the tube: stamens included : capsules subglobose, about 3 mm. in diameter, 
glabrous: seeds about 0.5 mm. long. 
In ponds and ditches, Massachusetts to New York, Florida and Louisiana. Spring and summer. 
2. SAMOLUS L. 
Annual or perennial caulescent herbs. Leaves alternate, various: blades entire. 
Flowers white or pink, in simple or panicled bracted or naked racemes. Sepals united at 
the base, persistent. Corolla perigynous: tube very short, the rounded lobes of the limb 
imbricated. Stamens 5, adnate to the corolla-tube, sometimes alternating with 5 narrow 
staminodia in the sinuses of the corolla-lobes. Anthers cordate, erect. Ovary 1-celled : 
stigma obtuse or capitate. Ovules numerous, half-anatropous, in a subglobose placenta. 
Capsule barely, if at all, elongated, 5-valved at the apex. Seeds numerous WATER 
PIMPERNEL. BROOKWEED. 
Racemes sessile or nearly so: staminodia at the sinuses of the corolla. 1. S. floribundus. 
Racemes long-peduncled : staminodia wanting. 
Peduncles glandular-pilose: corolla 4-5 mm. broad ; lobes broadly cuneate. 2. S. cuneatus. 
Peduneles glabrous: corolla 6-9 mm. broad; lobes suborbicular. x 
Glaucous: corolla 8-9 mm. broad. 3. S. alyssoides. 
Bright green : corolla 6-7 mm. broad. 4. S. ebracteatus. 
1l. Samolus floribündus H.B.K. Light green, glabrous, 1-6 dm. tall. Stems 
simple or diffuse: leaves few; blades obtuse, mainly 3-15 cm. long, spatulate or oval, 
or obovate, narrowed into winged petioles, the lower stem-leaves like the basal, passing 
through oval to ovate, the upper ones sessile or nearly so: racemes sessile or very short- 
peduncled : pedicels 1-2 cm. long, spreading or ascending: calyx campanulate ; lobes 
