242 MR. H. W. PUGSLEY : A REVISION OF THE 
than Cosson, and to retain Jéupicapnos as a genus. Its natural position in the 
family Fumariacer is between the genera Sarcocapnos and Fumaria, to both 
of which it approximates in certain features though possessed of other 
characters found in neither of them. 
CLASSIFICATION OF THE GENUS RUPICAPNOS. 
The genus Aupicapnos is a small group of plants of restricted geographical 
range. So far as can be judged from the sparse herbarium material 
available for examination, including, however, the important collection in 
Herb. Mus. Paris, kindly lent by M. Lecomte, it embraces under four well- 
marked sections a number of critical forms whose limitations are not always 
readily defined, just as happens with the better known and more widely dis- 
tributed genus Fumaria. Pomel,in his second account of the genus, divides 
his species into three subgroups, viz.: perennials with large flowers, perennials 
with small flowers, and small-flowered annuals. These form natural sub- 
divisions, unconnected, so far as is known, by intermediate forms, and have been 
adopted accordingly. The large-flowered species, with curved, purple-tipped 
corollas never very broadly winged, somewhat recall Fumaria agraria in 
their floral features (Pl. 9, figs. 4, 5). Their flowers are much more showy 
than in the other species of the genus, and the beauty of F. africana is 
commented on in Lamarck’s original description. It is proposed to place 
these plants in a section. Callianthos. 
Pomel’s second group, the small-flowered perennials, resembles the 
preceding section in habit and in fruit, albeit usually smaller in all its parts. 
Its flowers, however, are widely different, not only in size but in colour and 
form, and are quite unlike those of the Fumarie (Pl. 9, figs. 6, 7). Their 
colouring is uniformly whitish, with a greenish or yellowish suffusion about the 
apex, and the apical purple colouring of the inner petals is practically confined 
to their interior. The two outer petals are considerably dilated towards the 
apex, with spreading margins, and the inner petals are apically winged, some- 
times very broadly so. These wings of the inner petals spread horizontally 
between the patent margins or wings of the two outer petals in such a manner 
as to give to the flowers a triple winged aspect, and the name Tripteryæ 
therefore suggests itself for the section. 
In this second section X. sarcogapnoides, described by Cosson, is included 
by Pomel, with a note that the actual plant was unknown to him. An 
examination of Üosson's type in Herb. Mus. Paris shows that its flowers are 
quite unlike those of the other members of the section, the outer petals being 
conspicuously dilated towards the apex and much longer than the narrowly 
winged, purple-tipped inner petals. The corolla indeed resembles that of a 
Sarcocapnos in miniature, as stated by Cosson, and it seems necessary to place 
this plant in a separate section, for which the name Sarcocapnoides is 
proposed. 
