338 MR. H. W. PUGSLEY : A REVISION OF THE 
RUPICAPNOS. 
Rupicapnos, Pomel, Mat. Fl. Atlant. 16 (1860) ; Nouv. Mat. Fl. Atlant. i. 240 
(1874). Fumaria sect. Petrocapnos, Cosson & Durieu in Bull. Soc. Bot. 
France, ii. 305 (1855) ; Willkomm & Lange, Fl. Hisp. iii. 878 (1880) ; 
Cosson, Comp. FI. Atlant. ii. 80 (1883-1887). 
Plant: perennes caudice decumbente caulibusque brevissimis aut rarius annuw 
namv caule suberecto brevissime ramoso. Folia in speciebus perennibus 
pleraque subradicalia, in omnibus semper longe petiolata, foliolis breviter 
petiolatis vel subsessilibus 1-2-pinnatisectis vel inciso-flabellatis irregulariter 
pinnata (R. muricarid exceptá) ; segmenta secundaria decurrentia ; petioli 
nunquam cirrhosi. Inflorescentia pedicellis fructiferis gracilibus elongatis 
tandem deflexis racemoso-corymbiformis. Bracteæ pedicellis fructiferis 
multoties breviores. Corolla uni- vel inæeque bicalcarata ; petalerum ex- 
teriorum margines apicem versus in statu juvenili jam multo explicati ut in 
genere Sarcocapnos ; petalum superius subplanum potius quam semi-cylin- 
drieum, ealearatum, superne gibbum format qui apicem vix attingit et 
margines sepissime patentes in limbum latum interdum dilatatos nunquam 
purpureos habet sed non alas in jugum laterale productas ; petalum inferius 
inferne gibbum apicem vix attingentem marginibus patentibus cireundatum 
formans sepe basi ipsá breviter calearatum vel saccatum fit; petalorum 
interiorum nervus medius apicem versus sepe conspicue alatus est. Floris 
characteres reliqui utin genere Fumarid. Fructus monospermus, indeli.scens, 
nuciformis, semper mucronatus vel rostratus sed haud apice biforeolatus 
(sine rimis in mesocarpio) est ; exocarpium tuberculis rugosum ; endocarpium 
omnino ad mesocarpium adhwrens ; semen apice rotundatum vel paulo depressum, 
sine rugá longá infra mieropylam. 
Perennial, or more rarely annual plants, always with short stems, growing 
in rock-clefts. Inflorescence corymbiform with pedicels which lengthen in 
fruit and become deflexed so as to carry the seeds to the cavities of the rocks 
on which the planis grow. Lower petal often shortly spurred. Fruit 
without apical pits. 
In the perennial species of Rupicapnos the root is quickly elongated and 
the leaves are at first all radical, forming a close tuft. After the first year, 
the plant develops a thick, branched, decumbent rootstock, from which arise 
very short, branched stems, bearing long-petioled leaves and subterminal 
corymbiform racemes. The foliage is usually less decompound than in 
Fumaria, and the leaf-segments are much more shortly petioled and more 
decurrent, the secondary petioles never being cirrhose. The flower-buds of all 
the species resemble those of the genus Sarcocapnos rather than Fumaria, 
having well developed margins to the two outer petals, while the spur of the 
upper one is very small. This spur, as seen in perfectly developed flowers, 
varies greatly in different species and is of importance for affording specific 
