492 DR. MARIE STOPES ON BENNETTITES SCOTTII, 
of the bundle-sheath. It is a definite layer, clearly to be seen, though less 
thickened than the upper epidermis, and broken by gaps of what may be 
stomata as well as being very often obscured by the thick tuft of hairs 
attached to it. These hairs invest the lower side of the leaflets in such a 
closely packed mass that they form what at first sight appears to be a 
definite tissue (see z, text-fig. 3, and figs. 9 & 11, Pl. 20). The hair-like 
nature of the mass and some of their attachments to the lower epidermis can 
be seen in text-fig. 2, and fig. 11, Pl. 20. Here and there the basal attach- 
ments to the epidermis cells can be seen, and some hairs are cut obliquely, 
but most run in a direction parallel to the leaf, so that in the transverse 
section of the leaf they are also cut at right angles to their length and 
present themselves as more or less circular cells, The area between one 
Text-fig. 4. 
4. e. 
Detailed anatomy of central portion of leaf. See p. 496. 
leaflet and its neighbours is packed with these cells, which consequently 
together form much the appearance of a true tissue. This false-tissue of 
hairs is often about equal in area and shape to the leaflet to which it belongs, 
but in the middle region of the frond where the pinnules are more closely 
packed (PI. 20. fig. 12, and text-fig. 4) it is eliminated, or reduced to a very 
narrow zone. 
Cells which seem to me to be a corresponding mass of hair-cells seem to 
be interpreted by Wieland (1906, p. 83) as a “lower hypoderm,” a " heavy 
sclerenehyma region oceupying all the space below the bundles," and thai 
which he takes for a wavy transfusion tissue of a single row of cells seems 
to me to be really the lower epidermis. All the layer below this, as 
shown in the lower part of the upper half of his text-fig. p. 83, is really not 
sclerenchyma within the leaf, but is massed hairs outside it. The thickened, 
