440 LVII. MELASTOMACEE (HOOKER). 
one or two pores, rarely by slits, connective often produced below ae 
a pedicel, which is often tubercled at its base, or spurred before or i 
hind or both. Ovary 2- or more -celled (1-celled in Memecylon) 5 ra @ 
terminal, usually slender, stigma simple or capitate ; ovules ae e, 
anatropous, crowded on placentas that project from the inner ang ay 
the cells (whorled round a central placenta in Memecylon). I a 7 
loculicidal capsule included within the calyx-tube or a berry. See : 
usually minute, cochleate or angular (in Memecylon solitary and large); 
albumen 0; embryo terete or subglobose in all but the Memecylea 
which have a large radicle and convolute or consolidated coty eda 
Herbs shrubs or trees, sometimes epiphytic or climbing. Stem usua a 
4-angled; branches opposite. Leaves opposite, exstipulate, usually 3-9- 
nerved. Inflorescence various. 
A very large Tropical American order, less abundant in Asia, and comparatively rare 
in Africa. 
SuBoRDER I. MELASTOMES. 
Ovary 1- or more -celled; ovules numerous; placentas in the inner 
angles of the cells. Fruit many-seeded. Seeds minute; embryo 
minute, terete or subglobose. 
Trine 1, Osbeckieze.— Ovary altogether or partially free, its top convex. ate 
enclosed in the dry rarely»fleshy calyx-tube, crown convex or conical, usually hispid @ 
the tip. Seeds minute, cochleate. Inflorescence various. 
* Anthers equal or nearly so. 
Calyx glabrous. Anthers 8; connective 2-tubercled at the base . 1, NEROPHILA. 
Calyx hispid or bristly. Anthers 8 or 10; connective dilated and 
2-tubercled at the base, not lengthened below. Ovary hispid at 
NE a ee eg ey Onno 
Calyx glabrous. Anthers 8 or 10, broad ; connective not tubercled, 
elongated below and continuous with the filament. Ovary gla- 
ee, 
Calyx bristly. Anthers 8, narrow, pore large; connective short or 
produced and 2-tubercled at the base. Ovary crowned with 
bristles Pees uly iin, Se es ae a 
Calyx glabrous. Anthers 10, connective dilated and produced at 
ue, whee ii font ae 
** Anthers unequal ; connective of the larger usually much produced at the base. 
Anthers 10; not or very shortly produced ; 2-tubercled in front ; 
flowers capitate. Bracts large, persistent. . . . . . . . 6, TRISTEMMA. 
Anthers 8-10, connective of the larger produced at the base with 
long appendage in front. Inflorescence and bracts various . . 7. Dissortis. 
3. GuyontA. 
4, AnrHEROTOMA- 
Trine 2. Sonerilese.— Ovary more or less adnate to the calyx-tube; its top flat or 
depressed, angular. Capsule enclosed in the dry calyx-tube; crown broad, truncate, 
depressed, with 3-5 flat facets, which are often produced into erect scales. Seeds 
straight, angular, never cochleate. Inflorescence usually scorpioid, 
Anthers 10, very unequal, connective lengthened at the base, and 
appendaged in front . . ¢ 3 eee ee ce ees GB, APRIL See 
Anthers 10, sub-equal, connective lengthened at the base, and ap- 
pendaged in front. . . . <SORe ‘ * . 6 . 9 CaLvos. 
