THE THEEE rOEilS OF LTTHRUil SAT.ICAKIA. IvJJ 



does not enclose any anthers. The end of the pistil is generally 

 bent upwards at right angles. The six longer stamens, with their 

 pink filaments and green pollen, resemble in size of the grains 

 and in all respects the longer stamens of the mid-styled form, and 

 both correspond in length with the long-styled pistd. The six 

 shorter stamens, with their uncoloured iilaments and yellow pollen, 

 resemble in size of the grains and in all respects the longer 

 stamens of the long-styled form, and both correspond in length 

 with the mid-styled pistil. The capsules contain fewer seeds on 

 an average than in eitlior of the preceding forms, namely 83'5, and 

 they are considerably smaller in size. In this latter respect, but 

 not in number, there is a gradation parallel to that of the length 

 of tlie pistil, the long-styled having the largest, the mid-styled the 

 next in size, and the short-styled the smallest seed. 



From this description we see that there are three distinct 

 female organs, or rather females as they are borne on distinct indi- 

 viduals, diifering in the length and curvature of the style, in the 

 size of the stigma, and in the number and size of the seed. In the 

 three forms, taken together, there are thirty-six stamens or males, 

 and these can be divided into three sets of a dozen each, differing 

 from each other in length, curvature, and colour of the filaments, 

 in the size of the anthers, and especially in the colour and 

 diameter of the pollen-grains. Each of the three forms bcar,^ 

 half-a-dozcn of one kind of stamens and half-a-dozen of another 

 kind, but not all three kinds. The three kinds correspond in 

 length with the three pistils : the correspondence is always be- 

 tween half the stamens borne by two forms with the pistil of a 

 third form. These remarks apply to the structure, and not, as yet, 

 to the functions, of the reproductive organs. 



I ascertained the average number of seed by counting them in 

 eight fine selected capsides taken from plants of the three fonns 

 growing wild, and the result was, as we have seen, for the long- 

 styled (neglecting decimals) 93, mid-styled 132, and short-styled 

 83. I should not have trusted this result, but I had a number 

 of plants in my garden which, from their youth, did not yield the 

 full complement of seed, but they were of exactly the same age 

 and grew under exactly the same conditions, and were freely 

 visited by bees. I took six fine capsules from each, and found 

 the average to be for the long-styled 80, for the mid-styled 97, 

 and for the short-styled 61. Lastly, I made numerous artificial 

 unions, and, as may be seen in the following Tables, these gave 

 in the long-styled an average of 90 seeds, in the mid-styled 117, 



