EPIPHYLLUM. 187 
Hermann (Par. Botavus Prodr. Add. 2. 1689) first used the name Epiphyllum when 
he listed the name Epiphyllum americanum. Haworth credited the name also to Hermann 
in 1812 when he established the genus. 
KEY TO SPECIES. 
A. Perianth-tube 7 to 9 times as long as the limb... 0.0... oe cee ccccccececceceee. 1. E. phylianthus — 
Flowers 25 to 30 cm. long......... 00.0 c cece ce cece cece ence eceenceeeeece 2. E. oxypetalum 
Flowers 10 to 15 cm. long. 
Margins of joints crenate......... 0... cece ccc eee cee ence eceeeeeeues 3. E. pumilum 
Margins of joints undulate..........00 00000 eee ce eee cc eecueueucs 4. E. caudatum 
C. Joints deeply lobed. 
Joints 2 to 7 cm. broad. 
Lobes of joints spreading; outer perianth-segments lemon-yellow......... 5. E. darrahii 
Lobes of joints pointing forward; outer perianth-segments reddish yellow.. 6. E. anguliger 
Joints very large, up to 25 cm. broad........... 0. cece cece teen ceueaes 7. E. granditlobum 
CC. Joints crenate or nearly entire. 
D. Joints deeply crenate, thick; perianth-tube bearing foliaceous scales...... 8. E. crenatum 
DD. Joints low-crenate to nearly entire; perianth-tube without foliaceous scales. 
E. Sinui of the joint-margins very narrow; flowers up to 20 cm. broad; 
stamens yellow............ 0... .. 0. cece ee ee ee ee eee . E. macropterum — 
EE. Sinui of the joint-margins open; flowers 15 cm. wide or less; stamens 
not yellow. 
Ovary and fruit bearing linear scales....... 0.0.0... c ce eee eee eee 10. E. lepidocarpum — 
Ovary and fruit without linear scales. . 
Flowers 10 to 13. cm. long. ........ 0. ccc cece ee eee ee sence 11. E. pittiert — 
Flowers 15 to 28 cm. long. 
Flowers about 28 cm. long; style orange.................004- 12. E. guatemalense 
Flowers 15 to 25 cm. long; style white or pink. ; 
Joints very stiff... 2... ce cc ce ee eee eens 13. E. strictum 
Joints flexible or moderately stiff. ; 
Joints very large, up to 1 meter long and 12 cm. wide..... 14. E. stenopetalum 
Joints smaller, rarely ever 7 cm. wide. ; 
Joints shallowly crenate or subdentate; species of Costa 
DSS Cr: 15. E. cartagense 
Joints deeply crenate; species of Tobago, Trinidad, and . ; 
Veriezuela. 2... ec eee ee ee eee eeeees 16. E. hookeri 
a 1. Epiphyllum phyllanthus (Linnaeus) Haworth, Syn. Pl. Succ. 197. 1812. 
Cactus phyllanthus Linnaeus, Sp. Pl. 469. 1753. 
Opuntia phyllanthus Miller, Gard. Dict. ed. 8. No.9. 1768. 
Cereus phylianthus De Candolle, Prodr. 3: 469. 1828. 
Phyllocactus phyllanthus Link, Handb. Erkenn. Gewachse 2:11. 1831. ; 
Rhtpsalis macrocarpa Miquel, Bull. Sci. Phys. Nat. Néerl. 1838: 49. 1838 (in most part). 
Rhipsalis phyllanthus Schumann in Martius, Fl. Bras. 4?: 298. 1890 (in part). 
Hariota macrocarpa Kuntze, Rev. Gen. Pl. 1: 263. _1891. ; 
Phyllocactus phythanthus paraguayensis Weber, Dict. Hort. Bois 957. 1898. 
Phyllocactus phylianthus boliviensis Weber, Dict. Hort. Bois 957. 1898. 
Phyllocactus phyllanthus columbiensis Weber, Dict. Hort. Bois 957. I 898. 
Epiphyllum gaillardae Britton and Rose, Contr. U. S. Nat. Herb. 16: 240. 1913. 
Phyllocactus gaillardae Vaupel, Monatsschr. Kakteenk. 23: 87. 1913. 
Elongated and much branched; main branches narrow, terete or 3 or 4-angled, woody ; termi- 
nal joints elongated, terete or 3-angled below, usually flat or thin, rarely 3-winged, bright green with 
a purple margin, sometimes 7 cm. broad, obtuse, the margin coarsely serrate, the teeth obtuse; 
flower slender, 25 to 30 cm. long, the slender tube very much longer than the limb, green, t chm 
greenish or white, its segments narrow, 2 to 2.5 cm. long; scales on flower-tube few, minute, sprea ling; 
Style long, slender, pinkish (Schumann says white); filaments short ; stigma-lobes wort, white; fruit 
oblong, 7 to 9 cm. long, somewhat 8-ribbed, bright red; pulp white; seeds large, black, numerous. 
According to De Candolle, the flowers are nocturnal and odoriferous. 
Type locality: Brazil. 
Distribution: Panama to British Guiana, Bolivia, Peru, and Brazil. Recorded from 
Paraguay. 
