174 Rhodora [OCTOBER 
Moreover, certain plants of the first F, generation, including both 
whites and yellows, were chosen as the parents of a second filial 
generation, and their seeds which also resulted from pollination within 
the gynodioecious strain, were planted in the spring of 1912. So 
many of the plants of the 1912 cultures failed to flower the first year 
that it was necessary to wait until this year to report upon the second 
F, and the F; generations. 
It may be recalled that the first F; generation of 137 plants contained 
13 plants which could not be classified either as typical 1st form or 2d 
form hermaphrodites. Some of them did not flower; others were 
short-spiked gynomonoecious plants. They were kept in order to see 
what they would be like if they flowered in the second year. Eight 
of them died or again failed to flower; five of them flowered. 
Plant No. 7, which had abortive round spikes the first time it 
flowered, had long spikes about half of which were bifurcated the 
second season. The flowers were strictly pistillate, with the stamens 
reduced to mere rudiments. This plant, structurally the only purely 
pistillate plant of the entire culture, flowered so imperfectly in 1911, 
that it could only be referred to in the former paper (l. c. p. 206) as a 
possible exception to the rule that none of the progeny of the 2d form 
hermaphrodite approached the pistillate condition more closely than 
the mother itself. "This plant, either because it was sterile or because 
the season was so late when it flowered that no pollen was available, 
failed to set seed. It died after flowering. 
Plant No. 28 was short-spiked and gynomonoecious the first year. 
In the second year some of the spikes were as long as in the rest of the 
culture. The long spikes, which flowered first, had only 2d form 
flowers in the lower 3 of the spike and Ist form flowers above. The 
shortest spikes, which were the last to bloom, had only 1st form flowers. 
Between the extremes there were various transitions. 
Plants Nos. 41 and 53 did not flower the first year. In the second 
year they were typical 1st form hermaphrodites. 
Plant No. 63 did not flower the first year. The second season some 
of the spikes had first form flowers only. Other spikes were gyno- 
monoecious with the two flower types variously intermingled. 
These records show that plants which were short-spiked the first 
year did not maintain this character the second year. This fact 
throws grave doubt upon the taxonomic validity of Plantago lanceolata 
var. sphaerostachya, set apart by De Candolle on account of its short 
