1913] Dartlett, — Sex Forms in Plantago lanceolata 175 
spikes, and accords with the experience of Druce,! who cultivated a 
short-spiked plant and found that it lost its distinctive character in 
the second year. 
As a check upon the accuracy of the classification of the first Fi 
generation, 18 whites and yellows, including all the individuals chosen 
as seed parents of the F» generation, were retained until they flowered a 
second time. They were essentially alike in both years. The original 
mother plant of all the cultures has likewise held perfectly to the char- 
acters which it showed the first year. It has now flowered four times 
and has been divided into 6 plants. 
Four of the F; plants chosen as parents of an F, were yellows. Of 
these, two, Nos. 9 and 20 were altogether typical and quite indis- 
tinguishable from the mother plant, one, (No, 12) had the stamens of 
the same length and shape as the mother plant, but the anthers were 
slightly greener, and one, (No. 65), had the anthers just as in the 
mother plant, but the filaments were longer. The second form 
progeny of Nos. 9 and 20 reproduced the mother plant exactly. The 
2d form progeny of No. 12 (50 plants) were also typical yellows with 
the exception of one plant, the anthers of which were slightly greenish 
as in the mother. Since the mother plant itself was recorded as a 
typical yellow in its second flowering season, its variation in the first 
season from the typical yellow character may have been due to some 
environmental factor which likewise affected the solitary one of its 
progeny which resembled it. If, however, the variation toward 
greenish anthers indicated a stronger pistillate tendency in No. 12 
than in the other yellows, it is significant that the progeny of the 
greenish-stamened mother included a greater proportion of yellows 
than any other F;. The 2d form progeny of No. 65 showed a con- 
tinuous variation in the length of the filaments. In some plants 
they were as long as in the mother plant, in others as short as in any 
typical yellow, but the anthers of all were of the typical 2d form. 
If the long stamens of No. 65 indicated a weaker pistillate tendency 
than existed in the other yellows, the long-stamened character is no 
doubt to be correlated with the fact that the Fs included a higher 
proportion of whites than the progeny of any other yellow. 
1 Druce, G. Claridge: Plantago lanceolata var. sphaerostachya. Brit. Journ. Bot. 
xlix (1911) p. 235. “A plant of this, which I brought back from Jersey last year, 
retained its short spike during the year, but has this year developed spikes indis- 
tinguishable from the type, as Dillenius says it did in the Eltham Garden. (See Dill. 
Herb. 97.)” 
