TUM US, WEISS NO C 12, PAR 
1915]  Evans,— Notes on New England Hepaticae,— XII 119 
6. CALYPOGEIA PALUDOSA Warnst. Kryptogamenflora der Mark 
Brandenburg 2:1117. 1906. C. Trichomanis, var. tenuis Aust. Hep. 
Bor.-Amer. 74. 1873. Kantia Trichomanis, var. tenuis Underw.; 
A. Gray, Manual, Ed. 6, 713. 1890. Calypogeia tenuis Evans, 
RHopoma 9:69. 1907. According to the Vienna Rules the present 
species should bear the name C. paludosa, although Austin recognized 
the plant much earlier than Warnstorf and gave it the varietal name 
tenuis. When the writer raised Austin’s variety to specific rank he 
was unaware of Warnstorf's C. paludosa, published the preceding 
year in the supplement to his Kryptogamenflora. K. Müller con- 
siders C. paludosa a simple synonym of C. sphagnicola (Arn. & Perss.) 
Warnst. & Loeske, but Schiffner ! has recently shown that this view 
cannot be maintained and that C. paludosa is more closely allied to C. 
Trichomanis (L.) Corda than to C. sphagnicola. In separating it 
from the latter species he finds differences in both gametophyte and 
sporophyte. In the gametophyte he emphasizes the fact that the 
leaf-cells of C. paludosa are thin-walled throughout while those of C. 
sphagnicola are thicker and show distinct trigones. He adds that the 
plants in C. paludosa are larger than in C. sphagnicola, that the leaves 
and leaf-cells are larger, and that the underleaves are less deeply bifid 
and have broader lobes. 
In the sporophyte he brings out the following differences: in C. 
paludosa the cells of the outer layer of the'capsule wall are in eight 
longitudinal rows on each valve and show no traces of local thickenings 
in their walls, the cells of the inner layer are in sixteen longitudinal 
rows and show annular bands of thickening, while the spores measure 
12-14 u in diameter; in C. sphagnicola the cells of the outer layer are 
in fourteen to sixteen longitudinal rows and usually show rod-like or 
band-like thickenings of the radial walls (some of which extend over a 
portion of the outer tangential wall), the cells of the inner layer are in 
twenty-four longitudinal rows and show annular thickenings (much 
as in C. paludosa), while the spores measure only 9-10 u in diameter. 
In Schiffner's opinion it is the structure of the capsule which shows 
most clearly the close relationship of C. paludosa to C. Trichomanis, 
although he admits that the gametophytes of the two species some- 
times resemble each other very closely. He reserves the name C. 
Trichomanis for plants with a paroicous inflorescence, C. paludosa 
! Krit. Bemerk. über europ. Leberm. 13:8,9. 1914. 
