1917] Butters,— Studies in Ferns — Athyrium 201 
OrEGON: without locality, Elihu Hall no. 682 (f. Hilliz). 
WASHINGTON: Columbia River, 1841, Hinds; without locality, 
G. R. Vasey no. 47 (f. strictum), and no. 46; Ewell’s yard, Quiniault, 
June 25, 1902, H. S. Conard no. 154; Cheney, Mrs. Susan Tucker. 
BritisH Cotumsia: Selkirk region: Revelstoke, altitude 1600 ft. 
July 22, 1890, John Macoun (f. strictum), and July 3, 1905, C. H. Shaw 
no. 793; Glacier, altitude 3900 ft., August 8, 1909, Butters and Holway 
no. 475. 
Coast region: New Westminster, 1899, A. J. Hill, an extensive 
series of specimens illustrating different stages in the development of 
this fern, and including type material of Gilbert’s two forms. 
AuasKA: “Unalaschka, Dr. Mertens,” from Herb. Acad. Petrop., 
labelled in Ruprecht’s writing, “Athyrium Filix foemina var. aspi- 
dioides Ruprecht”; White Pass, July 23, 1914, Alice Eastwood no. 
874; Skagway, July 20, 1914, Alice Eastwood no. 796. 
4. Tue Lapy FERN or CALIFORNIA,! THE Basın REGION, AND THE 
SouTHERN Rocky MOUNTAINS. 
The lady fern of this region has been treated by all American authors 
as the true Athyrium Filix-femina. It is, however, found to differ 
from the typical form of that species in several minor respects, and 
in one important character. It should, therefore, be considered as a 
well marked geographical variety: 
ATHYRIUM FILIX-FEMINA (L.) Roth. var. californicum var. nov. 
differt a forma typica paleis atris, indusiis interdum dentatis 
vel brevissime ciliatis, sporis majoribus reticulatis nigrescentibus. 
The scales of this variety are usually considerably darker than in 
typical A. Filix-femina, but as in the case of similar dark scales found 
occasionally in European specimens, they have the same structure 
as the ordinary pale scales of A. Filix-femina, and not the fibrous 
structure characteristic of the dark scales of A. angustum. Short 
asplenioid sori are more frequent than in typical A. Filix-femina, and 
the indusia are seldom long ciliate, but rather short ciliate or merely 
toothed. In this respect they resemble those of A. angustum, but their 
dimensions are rather those of typical A. Filix-femina. They measure, 
on the average, 0.5 mm. high, and 0.8 mm. long. As in all American 
forms of A. Filix-temina, the sporangia are entirely without stalked 
glands. The most important peculiarity of this variety is the char- 
acter of the spores, which are unusually large, averaging 43 X 27.5 u, 
