1920]  Weatherby,—Varieties of Pityrogramma triangularis 117 
Those from the coastal region from Sonoma County south to Santa 
Barbara County, on the contrary, show no trace of these glands and 
average somewhat larger in size. "This difference, however, corre- 
lates with no other character and seems to call for no taxonomic 
recognition. Chamisso’s type collection of P. triangularis was made 
at San Francisco Bay and doubtless belongs to the glandless form. 
The original material of Gymnogramme oregana Nutt., preserved at 
the Philadelphia Academy, consists of two plants, one well-developed, 
with stout stipes 2-5 dm. long, the other small, with slender stipes 
1-2 dm. long, and of two detached fronds. In all, the fronds are 
very young, the lamina hardly fully expanded, and have a whitish 
indument. They show the red glands which seem to be character- 
istic of the northern material of P. triangularis. Nuttall himself 
has, on his label, crossed out the name oregana and written in triangu- 
laris with a reference to Hooker and Greville's plate of that species. 
Var. viscosa (Nutt. ex. D. C. Eaton), n. comb. Gymnogramme 
triangularis, var viscosa D. C. Eaton, Ferns of N. Am. ii. 16 (1879) 
and in Watson Bot. Cal. ii 335 (1880). Gymnogramme viscosa Nutt. 
in D. C. Eaton, Ferns of the Southwest 305, (1878) as syn. and 
Ferns of N. Am. l. c. Gymnogramme pyramidalis Nutt. in D. C. 
Eaton, 1. c., as syn. Ceropteris viscosa Underw. Bull. Torr. Bot. 
Club xxix. 631 (1902); Christensen, Ind. Fil. 170 (1905).  Pityro- 
gramma viscosa Maxon, Contr. Nat. Herb. xvii. 173 (1913). 
CALIFORNIA. SAN DigeGo County: Mission Hills, May 5, 1903, 
LeRoy Abrams, 3396 (G, S: glandular above; basal segments pinnat- 
ifid); shady ledges, Howard Cañon, La Jolla, Apr. 14, 1914, F. E. & E. 
S. Clements, 2 (G: indument yellowish); Rancho de la Nacion, 
Kimball (G); sides of ravines, Del Mar, March, 1894, Canby (G: 
glandular above; mixed with P. triangularis); dry hillsides, Linda 
Vista, July 6, 1915, Macbride & Payson, 788 (G); Jamul Valley, 
1875, Palmer, 433 (G: glandular above); Eucalyptus Cañon, hb. M. 
Rodman (G); moist ravine, 15 miles north of San Diego, March 7, 
1862, J. G. Cooper, 439 (N: glandular above); Otay Mesa, May 14, 
1915, Collins & Kempton, 79 (N); San Miguel Mts., Feb.-May, 
1900, Kimball (N: glandular above); San Diego, March 14, 1882, 
M. E. Jones, 3067 (N); Apr., 1875, hb. G. C. Woolson (Y); D. Cleve- 
land (Y); Pala, June 1880, Parish (Y); Evendido, Apr. 11, 1914, 
Parish, 9092 (S); Coast Mts., June, 1897, Parish (S: mixed with 
P. triangularis); Old Mission Dam, alt. 350 ft., Apr. 10, 1904, H. P. 
Chandler, 5055 (S: glandular above); San Diego, Nuttall (P: types 
of Gymnogramme viscosa and G. pyramidalis). RIVERSIDE County: 
Pigeon Pass Road, near summit of Box Springs Mt., alt. 1800 ft., 
Feb. 27, 1910, Reed, 2947 (N); Los ANGELES County: Santa 
