EPINANNOLENE, 107 
in this genus only a single pair of appendages, namely the posterior pair, of the seventh 
segment is retained, this appendage being, as in the other allied groups, the phallopod, 
and that its apparent double nature, suggesting its composition of two appendages, is 
due to its being secondarily biramous. This suggestion involves the supposition that 
the appendages of the first pair, the coleopods of other groups, are suppressed. There 
are difficulties in the way of the acceptance of both views of the matter, and the 
question at issue will probably remain unsettled until the development of the apparatus 
in question has been worked out. The view above advocated as to the apparatus being 
composed of a single pair of appendages appears to be that of Silvestri, who in his 
description of Physiostreptus says: ‘‘Organum copulativum pari uno appendicium 
constitutum ut in Pseudonannolene” (Boll. Mus. Torino, xviii. no. 433, p. 14, 1903). 
The two species of the genus Hpinannolene recorded below are very closely related. 
Analysis of the descriptions yields no very satisfactory characters for distinguishing 
them, apart from the structure of the copulatory apparatus of the males. By means 
of this organ they may be distinguished as follows :— 
a. Anterior branch of copulatory apparatus with the distal extremity bent and ending 
in two lamelliform processes, the external of which is denticulated on its inner 
edge 2. 6. ww ee ee eee ee eee ee Lpittieri.] 
a’. Anterior branch of copulatory apparatus evenly narrowed distally, not bent, and 
terminating in two subsimilar spiniform processes. . . . . . «© + + © « bicornis. 
[1. Epinannolene pittieri. 
Epinannolene pittiert, Brdlemann, Ann. Soc. Ent. France, Ixxii. p. 186, t. 1. figs. 3-7 (1903) °. 
Colour dark brownish-red, sometimes palely annulated; legs dark coloured. 
Body slender, shining. Head smooth; antenne short, hardly surpassing the first segment. Hyes narrow, 
consisting of three rows of ocelli. rst segment with its antero-inferior borders forming an evenly 
convex curve, marked with two strong grooves in addition to the marginal groove. Segments with their 
exposed portion very finely reticulated. The transverse sulcus distinct and marked with punctuations 
which are weak dorsally, stronger laterally ; longitudinal strie about five in nnmber and confined to the 
inferior portion of the segments. Sterna smooth. Anal segment moderately long; tergal plate with 
its rounded posterior border covering the valves, the edges of which are not compressed but are furnished 
with three pairs of setee. Sternal plate with its posterior edge transverse. 
Number of segments 49-55, 
Length 21-24 millim.; width 1-1-20. 
Hab. Cocos Isuanp (Biolley *). 
This species is included here for comparison with E. bicornis; the locality, however, 
does not come within the limits of this work. | 
2. Epinannolene bicornis. 
Epinannolene bicornis, Brélemann, Ann, Soc. Ent. France, lxxiv. pp. 856-358, t. 9. fig. 16 (1905) *. 
Colour dark blackish-brown, annulated ; legs yellowish. 
First segment with only one sulcus in addition to the marginal one. Exposed portion of segments without 
P2 
