1922] Fassett,—Lophotocarpus on River-Estuaries 71 
LOPHOTOCARPUS ON THE NORTH-EASTERN 
RIVER-ESTUARIES. 
NorMAN C. FassETT. 
(Plate 137.) 
IN Gray's Manual, edition 5 (1867), Engelmann described Sagit- 
taria calycina, growing from Maine to Delaware, Wisconsin, and 
southward, and added “var. spongiosa, with a loose or spongy tex- 
ture and linear bladeless leaves submersed, occurs eastward." In 
1894 J. G. Smith transferred S. calycina to the genus Lophotocarpus!, 
keeping var. spongiosa as a synonym of L. calycinus Smith. Later 
he described Engelmann's variety as a species’. 
H 
relied upon were as follows: 
L. calycinus. 
Leaves floating or ascending, 
[Leaves] 1.5—4 dm. high. 
Blades entire, hastate, sagittate, or 
triangular crescent-shaped, dorsal 
lobes usually widely divergent, 8- 
16 cm. Jong, 5-25 cm. wide, 
rounded, obtuse or acute at the 
apex. Basal lobes often much 
longer than the middle one. 
Scape shorter than the petioles, sim- 
ple, weak, at length decumbent. 
Verticils 2-6; fertile pedicels very 
thick, recurved in fruit, equaling 
or longer than the slender sterile 
ones. 
Achenes obcuneate, truncate, 2-2.5 
mm. long, narrowly winged on 
the margins, with a short, hori- 
zontal, triangular beak. 
1 Mem. Torr. Bot. Club, v. 25 (1894). 
The characters 
L. spongiosus. 
Submersed aquatic, with thick, spor- 
gy, nodose petioles and scapes. 
Petioles 1-2 dm. long. 
Blades spatulate and obtuse, or ellip- 
tical and truncate, or hastate, or 
sagittate, with the narrow, acute, 
falcately divergent lobes 2.5-10 
mm. wide, sometimes 8 cm. long. 
Scape simple; not more than half 
as long as the petioles, spongy, 
weak, at first ascending, at length 
decumbent, bearing two verticils of 
one to three flowers each. 
Fertile pedicels much thickened, 
1-2 or 3 cm. long, 3-5 or 6 mm. 
in diameter. 
Sepals broadly ovate, obtuse, scar- 
ious at the tip and margins, 10-12 
mm. long. 
Fruiting head depressed-globose, 7- 
10 mm. in diameter. 
Achene 2-2.5 mm. long, obcuneate, 
with a narrow dorsal wing and a 
very short, ascending or horizon- 
tal beak. 
2 Rev. of the Spec. of Loph. of the U. S., 4 (1899) and Rep. Mo. Bot. Gard. xi. 
148 (1900). 
3 Much of the material of L. calycinus has thick, spongy petioles, and nearly all 
of the petioles are nodose. 
4 Many apparently normal specimens in the Gray Herbarium have the leaves 
only 4 cm. long. 
