62 Rhodora [APRIL 
two thousand sheets of herbarium material, it was found that the 
nutlet-characters indicated in a striking way the relationships of 
species and sections within the genus. 
At this point it will be convenient to establish the generic and 
tribal relations of Scutellaria. In 1832-36, Bentham (Labiatarum 
Genera et Species) put Scutellaria and Perilomia—a South American 
genus—into a single tribe, but later, in 1848 (De Candolle’s Prodro- 
mus), he added Brunella and Cleonia to these two and threw them 
into a sub-tribe Scutellarieae of the tribe Stachydeae. Briquet, in his 
treatment of the genus in Engler & Prantl’s Die Natürlichen Pflanzen- 
familien, assigns Scutellaria and Salazaria Torrey to the tribe Scutel- 
larioideae, placing Perilomia in a separate tribe, Stachyoideae. It is 
noteworthy that in his grouping of the Perilomia entities he includes 
certain forms which other authors have put with Scutellaria, e. g., S. 
Mociniana Benth., and which apparently do not differ from that 
genus except by the alleged upright position of the seed. This special 
portion of the genus even has the scutellum on the calyx,—a character 
lacking in other sections. Undoubtedly the position of the seed is 
important in the classification, but its infallibility is questionable 
when it separates such apparently closely related forms. It is to 
be regretted that more material of these forms is not at hand. It is 
thought that the presence of the scutellum might serve as a more 
reliable generic character. 
Scutellaria then, is characterized by its bilabiate calyx, with lips 
entire and closed in fruit. From the upper lip is a projection known 
as the scutellum. Its co-genus Salazaria is set off on account of its 
calyx which becomes swollen in fruit. 
As somewhat detailed accounts of the genus, we have the earlier 
work of Arthur Hamilton, "Monographie du Genre Scutellairé,’’ 
Bulletin Seringe, 1832, and Bentham's treatise in the work above 
mentioned. Both these earlier writers have seen fit to make a number 
of sections, separated according to the nature of the inflorescence, 
Bentham confessing on his part the inadequacy of these characters. 
Hamilton makes but three sections and includes in them fifty-two 
species, fourteen of which are assigned to North America. Bentham 
makes five sections into which sixty-three species are placed, fifteen 
of which are North American. Inasmuch as in the present investi- 
gation the fruit has been specially studied, and since the Old World 
specimens available were too infrequently fruiting for accurate 
observation and too scarce for authoritative judgment, it was thought 
