348 COPELAND. 
true Goniophlchium. They have also a general aspect familiar in Ooniophlebium, 
but still more so in Eupolypodium, In Goniophlehium aimply pinnatifid fronds 
with the segments dilated at the base are much commoner in the American plants 
referred to this sub-genus, but occur also in a number of Ohl World species, 
especially in northern India. On geographical grounds, and because of the 
resemblance to Eupoly podium, and because ot less structural specialization, this 
would naturally be regarded as the primitive form of Goniophlehium, from which 
r 
"f!hcUolcpis" is a derived group. 
The primitive and generalized character of these plants is muclx more evident 
in the venation. In both species there is a costal row of areolse, but these areolw 
contain no free veins. The sori are borne sometimes on the vein which incloses 
the areola, sometimes against this vein but on a rudimentary branch excurrent 
from it. P. proavitum occasionally has other veins excurrent from this one. 
The venation then is neither that of typical Goniophlehium nor of Phymatodes, 
but is a generalized one. suggesting both of these. The structural similarity 
of Goniophlehium (Shellolepis) and Phymatodes and Setliguea I have already 
noted.^ Mr. ilaxon lias since told me that in his opinion these constitute a single 
gronp properly to be regarded as a genus. The discovery of these Bornean ferns 
is the best possible support for his judgment as to the affinity of Phymatodes 
(and tHelliyuea) to Goniophlehium, and shows that Goniophlehium, including 
them would certainly be a natural genus: but, so is Polypodium natural; and, 
unwieldy though it is, there is no hurry about di^niomberiiig it- 
4 
Polypodium sablanianum Christ Philip. Journ. Sci. 2 (1907) Bot. 177. 
Sarawak, Mount Bidi, ler/. Brooks. 
Frondibus pubescentibus. grandibiiSj herbaceis, sericbus areolanim 
praestantiiim duabus, a P. myhocarpo (Pr.) Mett. diversum. 
The fronds of the Bornean specimens are more than 00 cm long and about 
em broad. 
Described from Philippine specimens. 
Polypodium ceratophyllum Copel. spec. nova. (Plate VII.) 
Polypodiiim epiphyticum^ rhizomate repente, 1 mm crasso, paleis 
ferragineis 2.5 mm longis setifonnibus basibus peltatis vestito; stipite 
gracile^ brunneofulvo, articulate, ca. 8 cm alto sed supra medium lamina 
dcenrrente alato; fronde, ala decurrente exclusa, ca. 2.5 cm alta, ca. 4 
em lata, itei-um I)ifurcata, ramis late divergentibus, superioribus ca. 2 
mm latis, obtusis, coriaceis, glabris, margine angusta cartilaginea sub 
lente levissime incisa; venis occultis, laxe anastomosaiitibus ; soris super- 
ficialibus, utroque latere costae nigrae uniseriatis. 
Sarawak, Mount PoS, alt. 1,300 m, Foxicorthy 205. 
A species of the Microterus gronp, but with dichotomous fronds. On a sepa- 
rate rhizome are sterile fronds, broad and short and only once or twice forked; 
they may be fronds of an immature plant, or the sterile fronds of an adult. 
Syngramma angusta Copel. spec. nora. 
Rhizomate repente, lignoso, 2.5 mm crasso, paleis lanceolatis minutis 
obscuris coronato: stipitibus eonfertis, nisi ad baseos sparse paleaceas 
glabris, frondium sterilum 5-8 cm fertilium ca. 20 cm altis; fronde 
^This Journal, 2 (1907) Bot. 74. 
