THE DEHISCENCE OF ANTHERS BY APICAL PORES • 



195 



times almost porif orm slits, is an interesting form but hardly 

 merits consideration as one in which dehiscence is by apical 

 pores. 



Ardisia is a large genus of over 200 species much in need 

 of revision. The stamens are 5 in number, with short 

 sagittate or lanceolate anthers usually opening by longitu- 

 dinal slits. In the section Icacorea, however, according to 

 Pax, the anthers open by apical pores. This section seems 

 to comprise about 7 South American species. The anthers 

 of the second section, Stylogyne, with 1 Brazilian and 1 



Malayan 



species, are 



described as opening by terminal 



pores. In the single representative of the third section, 

 A. paludosa of Madagascar, the anthers open by a single 

 terminal pore. In the two other sections of the genus 

 containing the remainder of the species, distributed 

 throughput the tropics of both hemispheres, dehiscence is 

 by lateral slits. My knowledge of this genus is very im- 

 perfect, but I think it probable that an examination of 

 suitable material would show that in the most of the above 



M 



species described as apically dehiscent the pores are not 

 permanent, but merely the 



beginnings 



Of 



longitudinal 



slits. 



Loganiaceae. 



In the 32 genera assigned to the Loganiaceae only 5 or 6 

 have any species with a rotate corolla, the tendency being 

 towards tubular, salver-form or campanulate. Gardneria 

 is the only genus characterized by a rotate corolla with 

 oblong to linear exserted anthers. Three species are 

 found in Japan and India. In G. nutans (of Japan) the 

 slits are said not to extend to the base as in the others, but 

 to be confined to the tip of the anther. 



Gentianaceae. 



In the Gentianaceae, two genera belong clearly to this 

 type: Exacum (29 species, distributed over tropical and 

 sub-tropical Asia, the Malay Archipelago, Madagascar and 





