THE DEHISCENCE OF ANTHERS BY APICAL PORES. 201 



particular subdivision, but in our almost total ignorance of 

 their ecology and in some cases of their structure it seems 

 best to treat these forms in this way. 



These so-called aberrant forms are the following : 

 Odontostomum (1 species in California) of the Liliaceae 

 has a hypocraterif orm corolla with patent or reflexed lobes 

 with erect, basifixed, ovoid anthers which are only about 

 one-half or one-third as long as the filaments. 



Poranthera ( 



o 



of which one also occurs in Tasmania and New Zealand, 

 and a sixth endemic in New Zealand) has small flowers with 

 five stamens having 4-locellate anthers opening in four 



sometimes 



fluent into two. 



In the Ebenaceae, the anthers of the three species of the 



m mm a m 



rluin (2 in Madag 

 ) of Diospyros ( 



) 



described as opening by lateral pores at the tip instead of 

 by longitudinal slits as in the other species. The corolla 

 in Diospyros is urceolate, campanulate, tubular, or salver- 

 form ; stamens 4 to indefinite, usually 16 ; anthers oblong, 

 linear or lanceolate. In the three species which have been 

 described as apically dehiscent the flowers are small, the 

 corolla seems to be openly campanulate and the stamens 

 10-16 in number. It may be that these species might be 

 properly assigned to the Solanum-Cassia type. Sargent 

 describes the sixteen anthers of D. Texana included in the 

 urceolate corolla as dehiscing only near the apex and his 

 figure represents the openings as almost poriform. 



°The condition prevailing in the Acanthaceae is one of par- 

 ticular interest. Here we have 7 genera which are said 

 to have, in some species at least, anthers opening by apical 

 pores. Of these, Staurogyne need not be considered, since 

 dehiscence in this genus can hardly be justly described as 

 by apical pores. Ophiorrhiziphyllon (1 species in Marta- 



