208 The Philippine Journal of Science 1922 
It may be distinguished from still another similar genus, 
Campbellosphaera, described by Shaw (719), by the absence of 
a vacancy in the layer of somatic cells of a mother over the 
gonidia and the daughters developed from the gonidia. In 
that genus the gonidia are not formed from cells in the coenobium 
wall at the places where they are to grow and develop into 
daughters, but come from the outside of the coenobium to take 
up places within. In the species which forms the subject of 
this paper the gonidia are differentiated from the somatic cells 
late in the prenatal development of the asexual daughters in 
which they are formed. As they grow they sink below the 
level of the somatic protoplasts, leaving a vacancy in the layer 
of protoplasts. 
This new species will be treated as the type of a new genus, 
to be called Copelandosphaera* as a token of recognition of the 
work of Edwin Bingham Copeland on the phylogeny of the 
ferns of the Oriental Tropics. The first specimens observed 
were living, and some of them appeared to be setting free their 
vegetative cells or somatic protoplasts, whence the name dissi- 
patrix that will be applied to the species. The use of this name 
is not intended to imply that the behavior to which it refers is 
habitual; nevertheless, it may serve to direct attention to a 
possible recurrence of the phenomenon in this or other species 
of the family under circumstances that may admit of following 
the history of the detached vegetative cells. 
COPELANDOSPHAERA DISSIPATRIX GEN. ET SP. NOV. 
DESCRIPTION OF TYPE SPECIMEN 
For the type of Copelandosphaera dissipatrix a mature asexual 
specimen, represented by Plate 2, fig. 6, and Plate 3, fig. 18, 
has been selected. It is in a Venetian turpentine mount of 
material that was collected at Pasig, near Manila, in August, 
1914. The collection from which the slide was prepared was 
numbered XVI.2- The material was stained with a combination 
of nigrosin and Bismarck brown. 
*The use of this name was forecast in a footnote of an earlier paper 
(Shaw 719, p. 513). 
* The slide bearing the type specimen is in my possession. Slide mounts 
of material collected not far from the type locality in the same year have 
been sent to Prof. F. G. Haughwout, Bureau of Science, Manila, P.‘I., and to 
Prof. D. H. Campbell, Stanford University, California. Material from the 
same locality, bottled in glycerine, has been sent to thirty-two biologists 
in the Northern Hemisphere. Duplicates of this bottled material are 
available for distribution from my American address: Claremont, Cali- 
fornia. 
