47,5 Merrill: Studies on Philippine Rubiaceae, IV 485 
31310, 31315 Ramos & Edafo, April and May, 1918, in damp 
forests along small streams. 
This species is readily distinguished from the others described 
in the genus by its indumentum and especially by its densely 
villous bracteoles and flowers. The bracteoles are deeply 4-cleft, 
the inner series with free or nearly free lobes. It has much 
the appearance of certain forms of Urophyllum but has the 
involucriform double series of bracteoles characteristic of 
Williamsia. 
WILLIAMSIA LONGISTIPULA sp. nov. 
Frutex erectus, plus minusve ciliato-villosus; foliis membra- 
naceis, oblongo-lanceolatis, usque ad 27 cm longis, supra glabris, 
tenuiter caudato-acuminatis, nervis utrinque circiter 17, per- 
spicuis, subtus ciliato-villosis; floribus axillaribus, fasciculatis, 
villosis, bracteolis involucrantibus membranaceis, villosis, ex- 
terioribus 3 mm diametro, profunde 4-lobatis, interioribus sub- 
quadratis, 4 ad 5 mm latis, angulis leviter acuminatis; fructibus 
ovoideis, villosis, circiter 8 mm longis. 
A shrub, 2 to 3 m high, the branchlets, stipules, flowers and 
leaves on the midrib and nerves beneath ciliate-villous, the older 
branches glabrous or nearly so, the internodes 3 to 6 cm long. 
Leaves membranaceous, oblong-lanceolate, pale when dry, 23 to 
27 cm long, 6 to 7.5 cm wide, the upper surface entirely glabrous, 
base acute, apex slenderly caudate-acuminate; lateral nerves 
about 17 on each side of the midrib, prominent on the lower 
surface, curved, ascending, arched-anastomosing close to the 
margin, the reticulations rather lax, distinct, these sparingly 
ciliate-villous as are the nerves and midrib; petioles 1 to 1.5 cm 
long; stipules narrowly lanceolate, 3 to 4 cm long. Flowers axil- 
lary, fascicled, villous, the basal, cuplike bracteoles membrana- 
ceous, villous, the outer one about 3 mm in diameter, deeply 
4-lobed, the lobes lanceolate-acuminate, the inner one nearly 
square, 4 to 5 mm wide, the corners acuminate. Fruits ovoid, 
about 6-celled, villous, about 8 mm long, sessile or nearly so, the 
persistent calyx lobes 4, broadly ovate, rounded or obtuse, about 
3mm long. 
MINDANAO, Butuan Subprovince, Agusan River at Waloe, 
Merrill 7287, October 2, 1910, in damp forests, altitude about 
50 meters. 
This species is most closely allied to Williamsia panayensis 
Merr., from which it is distinguished by its longer stipules, its 
en leaves, and by its entirely different involucral 
racts. 
