﻿PEDALINEiE (Stapf). 455 



at the base of the back ; limb obscurely 2-labiate, usually short. 

 Stamens 4, more or less distinctly didynamous, with the rudiment of 

 the fifth present (very rarely 2 fertile and 2 staminodes), inserted 

 and enclosed in the corolla-tube, rarely shortly exserfced ; anther- 

 cells 2, dehiscing longitudinally, hanging from the apex of the 

 connective and often somewhat divergent, or dorsally attached to it 

 and parallel, connective nearly always with an apical gland. Hypo- 

 gijnous disc always more or less developed, often asymmetrical. 

 Ovary superior, very rarely inferior, sessile, 2-4- (rarely 1-) celled ; 

 cells often completely or incompletely divided by spurious septa ; 

 style filiform, slightly exceeding the anthers ; stigma 2-lobed, lobes 

 ovate to linear ; placentas central; ovules 1 to many in each cell. 

 Fruit very variable, dehiscent or indehiscent, often provided with 

 spines, horns or wings. Seeds 1 to many in each cell, sometimes 

 winged, with a delicate or stout testa; albumen very thin; embryo 

 straight, cotyledons flat, radicles short. 



Annual or perennial herbs, rarely shrubs or small trees, more or less covered 

 with sessile mucilage glands (at least when young) ; leaves opposite or the upper 

 alternate ; flowers mostly axillary and solitary, rarely in few- to many-flowered 

 axillary and terminal inflorescences: pedicels usually with ncctarial glands 

 (modified flower-buds) at the base. 



DiSTEiB. Species about 55, in the tropics and the extra- tropical countries of the 

 southern hemisphere of the Old World. 



All the African Pedalineee belong to the tribe Pedaliece. 



I. Pterodiscus.— CoroZ/a funnel-shaped, oblique or slightly gibbous at the 

 base. Ovary and fruit 2-celled ; cells undivided, 1-2-ovuled. Fruit inde- 

 hiscent, winged. 



II. KolvLbia.— -Corolla funnel-shaped from an oblique, widely saccate base. 

 Ovary 2-celled ; cells undivided ; 8 ovules in each cell. Fruit unknown. 



III. Harpagophytum.— Corona funnel-shaped, equal or slightly gibbous at 

 the base. Ooary and fruit 2-celled; cells undivided; ovules numerous, 

 2-seriate. Fruit tardily dehiscent, armed with hook-bearing horns. 



IV. Roseri&. —Corolla funnel-shaped to subcylindric, subgibbous or subsaccato 

 at the base. Ovary and fruit very unequally 2-cellod; cells mcompletely 

 divided by false septa. Fruit with 2-8 unequal spines or tubercles at the 

 base; posticous cell small and indeliiscent ; auticous cell many-seeded, 



loculicidal. i. n t 



V. Seammum. — Corolla obliquely campanulate, obscurely gibbous at the ttase. 

 Ooary and fruit equally 2-celled; cells incompletely divided by false 

 septa. Fruit a many-seeded loculicidal, acute or beaked, unappendaged 

 cuusuIg 



VI. Ceratotheca.-Like Sesanuim, but the capsule with 2 divergent horns or 



spines at the apex. , ^ i / •* 



VII. Pretrea.-Co)-oi?a pendulous, obliquely campanulate. oar j/ and J rutt 

 2-celled, cells completely divided by false septa, with 2 ovules lu each division. 

 Fruit indehiscent, disc-shaped with 2 central spines. 



I. PTERODISCUS, Hook. 



Calyx small, 5-partite. Corolla: inhe f;^.°^^«l-^haped oblique or 

 slightly gibbous at the base; limb spreading, aubbilabiate, lobes 

 orbiculart subequal. Stamens didynamous; anthers converging; 



