249 
Appendages to the Petioles of Liriodendra, 
By ARTHUR HOLLICK. 
(PLATES 269, 270.) 
In a paper read some time ago before the Club and published 
in the BuLLeTin* attention was called to certain wing-like ap- 
pendages on the petioles of fossil leaves considered to be allied to 
the living Latodendron Tulipifera L. Referring to the theory ad- 
vanced by Professor Lester F. Ward in regard to stipular appen- 
dages to the petioles of Platanus,} that they may represent former 
basal lobes of the leaf, which have become detached, forming first 
basilar appendages, then wings to the petioles, and finally stipules, 
I suggested that we might have a similar example in the case of 
Liriodendron. In other words, that the stipules of the living 
Species had been derived from the wing-like appendages, which in 
turn had been previously derived from basal leaf lobes. 
Since then a series of leaf forms of Liriodendron was sent to 
me by Mr. Edward W. Berry, of Passaic, N. J. They were col- 
lected from saplings, seedlings and new shoots from old stumps, 
and many of them are exceedingly interesting and significant. In 
the specimen to which I wish to call special attention the blade. 
of the leaf is abnormally broad at the top and narrow at the base. 
The petiole is short and has broadly winged margins, which ex- 
tend from the base of the petiole and connect with the base of 
the leaf blade. On one side there is a sharp sinus where they 
join, on the other merely a contraction. The nervation of each 
extends into the other. The question to decide is whether we 
have stipules adnate to the petiole and leaf blade, or portions of 
the leaf blade which are acting the part of stipular appendages. 
The abnormal shape of the leaf is also significent, suggesting 
at once the fossil genus Liriophyl/um, in which at least one species 
(L. populoides Lesq.) is known to possess winged petioles. 
On plate 269, for purposes of comparison with our specimen, 
Thave introduced a figure of this species and one which shows 
*Bull. Torr. Bot. Club, 21: 467-471. p/. 220,221. 1894. 
}Proc. U. S, Nat. Mus. 11: 39-42. p/. 17-22. 1888. Am, Nat. 24: 797. 810. 
Pt. 28. 1890, 
