RYDBERG: SOME GENERIC SEGREGATIONS 279 
Corolla scarcely 2-lipped ; almost equally deeply 4-lobed. 
Corolla tubular-funnelform; capsule neither flattened nor obcordate at the 
apex, 4-valved. 
Leaves alternate but mostly basal. Wulfenia. 
Leaves opposite; stem leafy. Leptandra. 
Corolla rotate or short-campanulate ; capsule flattened, obcordate, 2-valved. 
Leaves opposite or whorled ; stem leafy. Veronica. 
Leaves alternate, but mostly basal ; plant with a mostly naked scape, 
Synthyris. 
Corolla, if present, deeply 2-lipped; upper lip entire, broad, arcuate ; lower lip 
straight and deeply laciniate ; leaves alternate but mostly basal ; scape bracted. 
Besseya. 
SyntTHyrkis Benth. ; DC. Prodr. 10: 454. 1846 
This genus was based on S. reniformis. S. rubra was also 
included in the genus by the author, but he admits that it is a 
rather anomalous member, lacking the corolla. It is, however, 
more nearly related to S. alpina, which has a corolla of a different 
structure. The species belonging here are the following : S. rent- 
formis (Dougl.) Benth., S. major (Hook.) Heller, S. prmnatifida 
S. Wats. and S. /aciniata (Gray) Rydb. Whether S. rotundifolia 
A. Gray should also be included or should be made the type of 
a separate genus is a little doubtful. At present Iam inclined to 
regard it asa Synthyris. Soalso S. schizantha Piper, with lacerate 
corolla-lobes, which species is still more aberrant. 
Besseya gen. nov. 
Low perennials with a thick rootstock. Basal leaves petioled ; 
blades ovate or oblong, crenate; stem-leaves bract-like, alternate: 
inflorescence a bracted spike: calyx almost regularly 4-cleft to 
near the base or cleft to the base on the upper side and then 2—3- 
lobed : corolla 2-lipped, cleft to near the base, or lacking; upper 
lip, if present, entire, obovate or cuneate-obovate, concave ; lower 
lip much shorter, often irregularly 2-3-cleft or laciniate : stamens 
2, attached at the base of the corolla, or if this is absent ona 
small disk: capsule flattened, obcordate, 2-valved. 
It is with pleasure that I dedicate this genus of western moun- 
tain plants to Professor Charles E. Bessey, of the University of 
Nebraska, my former teacher and the most prominent botanist of 
the region where these small plants grow. 
The first species cited below is regarded as the type. 
