450 Nasu: REVISION OF THE FAMILY FOUQUIERIACEAE 
ovary, the more or less united styles and the gamopetalous tubu- 
lar corolla, to the base of which the filaments are slightly adnate, 
all markedly point in this direction. The spongy central column 
found in the dehiscing capsules would indicate that the inner edges 
of the septa unite. A transverse section of the ovary made at this 
time, and before the septa break away from the walls, would much 
resemble the condition of things existing in the genus Gz/za, also 
found in the same region. The general resemblance of the flow- 
ers to those of some of the Yarge-flowered red Gilias is perhaps the 
most striking feature of the plants. It is true that the sepals in 
this are distinct, while in Gia they are more or less united, but 
this is but one character weighing against many others. It would 
seem to me that the ovarian, placental, style and corolla charac- 
ters indicate a strong affinity with that family, much stronger than 
with the Tamaricaceae, to which family, polypetalous in all its 
other forms as are its immediate relatives, the gamopetalous corolla 
of Fouquieriaceae does violence. 
FOUQUIERIACEAE DC. Prod. 3: 349. 1828. 
Shrubs or trees, the trunks simple, columnar and stout, or 
much branched: branches spine-branching. Spines developed 
within the petioles of the leaves on the new growth and becoming 
apparent when these fall. Leaves with the blades flat, entire, or 
rarely obcordate or emarginate: those on the young growth 
petioled, the petioles from one half as long as to equalling the 
blade. Secondary leaves borne in fascicles in the axils of the 
spines and sessile or nearly so. Inflorescence spicate to paniculate, 
terminal. Flowers sessile or pedicelled. Calyx of five free im- 
bricated sepals. Corolla yellow or red, hypogynous, the segments 
united for one half their length or more: tube cylindric, some- 
times broadened toward the apex: lobes flat or concave, imbri- 
cated in the bud, incurved, erect, spreading, reflexed or enrolled. 
Stamens ten to fifteen, of unequal length, exserted from the corolla- 
tube, and adnate to it at the very base : filaments subulate, usually 
broadened and compressed dorsally at the more or less pubescent 
base where they are sometimes coherent, sometimes provided 
with a scale-like appendage near the base: anthers acute at the 
apex, cordate at the base, elliptic to nearly ovate, introrse, versa- 
tile, attached below the middle, 2-celled, the cells opening longi- 
tudinally. Ovary 3-celled, the inner edges of the septa united at 
the base and at the very apex, free in middle, the septa finally 
