Hepaticae of Puerto Rico 
Il, HARPALEJEUNEA, GYRTOLEJEUNEA, EUOSMOLEJEUNEA AND TRACHYLEJEUNEA 
By ALEXANDER W. EVANS 
(WITH PLATES 20-22) 
HARPALEJEUNEA 
The subgenus Harpa-Lejeunea as first limited by Spruce 
included three sections.* In the first of these, Cardiostipa, the 
underleaves were described as cuneate-obcordate or cuneate-bifid 
with very obtuse angles or segments ; while in the second, Cyc/o- 
stipa, they were said to be suborbiculate and, with a single excep- 
tion, bifid to the middle with acute or very rarely obtuse segments. 
The exception was Lejeunea holostipa Spruce, a species with 
undivided underleaves. The third section, Strepsi-Lejeunea, was 
made up of more robust forms with strongly thickened cell-walls 
and much larger underleaves. Spruce implied at the beginning 
that this third section was almost of subgeneric value and a few 
years afterwards wrote of it as a subgenus. When Schiffner ele- 
vated Spruce’s subgenera to generic rank, he made two genera, 
tentatively at least, out of the subgenus Harpa-Lejeunea as origi- 
nally defined. For the first of these, which still included the 
sections Cardiostipa and Cyclostipa of Spruce, he retained the 
name Harpalejeunea. To the second he naturally gave the name 
Strepsilejeunea. Both of these genera are fully recognized by 
Stephani.{ 
The sections Cardiostipa and Cyclostipa are really of unequal 
value. Cardiostipa is a very natural group, characterized espe- 
cially by a peculiar type of underleaf, while Cyclostipa is made up 
of rather heterogeneous elements. This fact was clearly recog- 
nized by Spruce, who stated that his three species, L. holostipa, L. 
* Hep, Amaz. et And. 164. 1884. 
+ Engler & Prantl, Nat. Pflanzenfam. 13: 126, 127. 1893. 
t Hedwigia, 35: 97, 127. 1896. 
544 
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SS cee eeteraiereaticpeeemrnntene 
