98 Evans : Genus of Hepaticae from Hawaiian Islands 



species of Bazzania, Pleurozia and Anastrophylhim. Both male 

 and female flowers occur somewhat abundantly and a few of the 

 latter show well developed perianths. Mature capsules, unfortu- 

 nately, are not present. The descriptions in the present paper are 

 drawn almost entirely from Mr. Cooke's material, which has also 

 served for the drawings. 



At a cursory glance, the plant looks unlike atypical Bazzania ; 

 in the first place because its leaves are transversely inserted and 

 strongly squarrose instead of being incubous and appressed, 

 and in the second place, because its underleaves are relatively 

 larger than those of Bazzania and are also strongly squarrose. 

 These peculiarities might seem to indicate an affinity with certain 

 genera of the Ptilidioideae, such as Mastigophora or Herberta or 

 the curious and imperfectly understood Herpocladium of Mitten. 

 The position of the sexual organs, however, both male and female, 

 on short branches axillary to the underleaves, shows at once that 



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the true affinities of the new genus are with the Trigonantheae. 



* 



The bracts and bracteoles, moreover, as well as the fully developed 

 perianths, are essentially like those of Bazzania and of certain other 

 genera of this same tribe. 



More important as a generic character than the foliar differences 

 just indicated is a peculiar kind of branching, unlike anything 

 hitherto described for the Hepaticae. In the related Bazzania, as 

 is well known from the studies of Leitgeb, the branching is of 

 two kinds. In the first, the so-called " Endverzweigung " or 

 11 terminal branching/'* the branches are exogenous in origin and 

 lateral, each branch representing the postical half of one of the 

 lateral segments of the apical cell.f The leaf, which normally de- 

 velops from the whole of such a- segment, is here restricted to the 

 antical half and is therefore narrower than an ordinary leaf and not 

 toothed at the apex. A branch here, which is always an ordinary 

 leafy branch, is of about the same size as the axis from which it 

 springs ; and, as the axis is itself deflected in the ooposite direction 

 from that taken by the branch, the plants of many species appear 



*Untersuch. iiber die Lebermoose, 2 : 22, 23. pL 4. f. 2, 4. 1875. 



f Branching from the postical half of a lateral segment occurs in many genera of 

 the Jungermanniaceee and (with the modification seen in Radula, etc.) is the only type 

 of terminal branching described by Leitgeb. 



