Durand : Classification of the Fleshy Pezizineae 477 



pacted parallel threads running in a longitudinal direction. As 



the cup expands at the top of the stem these hyphae spread out in 

 a radiating manner toward the margin. At the side of the hy- 

 menium they expand, forming a long and prominent margin to the 

 cup. At this place they become thicker and looser, with frequent 

 septa, thus forming a strongly pseudo-parenchymatous tissue, with 

 cells 6-10 fi in diameter. The marginal pili are direct prolonga- 

 tions of these hyphal cells. At the surface of the cup these hy- 

 phae are replaced by a thin stratum of pseudo-parenchymatous 

 cells, 5-6 fi in diameter. 



S. coccinea (Scop.) Sacc. PL 28, Fig. 4 



The hypothecium in this species cannot be well distinguished 

 from the adjacent excipular tissues. The excipulum consists of 

 two well-marked layers. The inner or medullary is very thick 

 and is composed of long, rather slender, hyaline hyphae loosely 

 interwoven. At the sides of the hymenium these become parallel 

 and closely compacted, with frequent septa, thus forming a wide 

 margin of cylindrical parenchymatous cells. This marginal tissue 

 closely resembles that described for S. floccosa preceding. 



The outer layer is composed of thin-walled pseudo-parenchym- 

 atous cells, 6 x 1 5-20 fi in diameter, which are elongated in a 

 direction parallel to the surface of the cup. Aside from being 

 different in form these ectal cells take a deeper stain than the 

 medullary hyphae. This layer in some cases separates quite easily 

 from the one within. 



The structure of the medullary tissue of Sarcoscypha coccinea 

 and S. floccosa suggests the prosenchyma found in the Helotiaceous 

 plants. In 1893, Schroter * included the genus Sarcoscypha with 

 part of Macropodia in the family Helotiaceae, remarking that it ap- 

 proaches closely to the genus Pcziza {Macropodia), referring es- 

 pecially to P. macropus. But the gross characters of these plants, 

 such as the size, form, and consistency of the cups associate them 

 more closely with the Pezizas, and other writers, including Rehm |, 

 have placed them among the Pezizaceous genera. It was diffi- 

 cult, therefore, before a study of the characters of the sterile layers 

 was undertaken to believe that these forms should be consistently 

 placed in different families. 



fRehm, p. 1070. * Schr.Her (1), p. 59. 



