520 - WHITE: THE SALTATORY 
peared which Professor de Vries believed had originated then and 
there by mutation from O. Lamarckiana. He transferred many 
of those plants and much of their seed to his experimental gardens 
and there verified that belief by obtaining a repetition of those two 
mutations under his direct observation. Furthermore, in the course 
of a few years of experimentation, he obtained by mutation from 
O. Lamarckiana five other new species, seven in all. A part of 
these new species proved to be also mutative, adding still other 
new ones to his list ; and all the new ones possessed, from the mo- 
ment of their origin, well-defined and heritable attributes. These 
mutations occurred in the progeny of plants which had received 
intraspecific artificial pollination. Interspecific pollination, or cross- 
fertilization, was not found to facilitate mutation, and hybridization 
was not sought for or desired. 
In view of the success of Professor de Vries’ experiments with 
Oenothera, and the fact that all except one or two per cent. of the 
species with which he experimented failed to show any sign of 
mutation; and the further fact that he experimented with only a 
part of the Holland flora, one infers that the proportion of immu- 
tative plants to the mutative may perhaps be as great throughout 
the plant kingdom as he found it to be in that flora. 
Recognizing the fundamental accordance of my observations 
with those of Professor de Vries, I assume that Lycopersicum 's, 
like Oenothera, in its mutative period and that L. esculentum has 
given mutative origin to two other species which, in previous para- 
graphs, I provisionally designated as groups. The fact that these 
two specific forms differ less from each other, and from their com 
mon parent ZL. esculentum, than do some other species from one 
another, does not affect the validity of the claim that they should 
also be regarded as distinct species. The facility with which all 
forms of Lycopersicum hybridize with one another also does not 
affect that claim because of the non-phylogenetic character * 
hybridization. I shall therefore apply the systematic aa . 
Lycopersicum solanopsis to the solanoid group, and the Jatifoliate 
group accordingly becomes L. datifoliatum ; but the atavic grouP 
must retain its original name of L. esculentum. : 
While the cases of mutation which I have observed ie 
esculentum, fundamentally accord with those which ProfessoF 
