450 Lloyd : Anatomy of Chrysoma pauciflosculosa 



By applying glycerine to thick sections of alcoholic material he 

 has in the majority of cases been able to produce the movement. 

 The results, however, are not convincing, and more exact studies 

 should be carried out upon living material. If this explanation 

 holds good, the position of the palisade cells may be accounted 

 for as contributive to the formation of the diaphragm. 



An alternative explanation may be found in the light relations. 

 Vertically placed leaves or their physiological equivalents are com- 

 monly supplied with palisade tissue on both faces (isolateral), but 

 hitherto no such peculiar grouping of the chlorenchyma in an 

 isolateral leaf has come to light Such grouping therefore ap- 

 pears to be secondary and to be accounted for in some other way. 

 The best we can do, therefore, is to admit our ignorance and 

 await experimental evidence. 



No further matters in the anatomy of this plant need receive 

 mention at this time beyond the remark that the shrubby character 

 of the plant and its peculiar leaf anatomy clearly separate it from 

 the genus Solidago, in which, so far as at present known, no ap- 



m 



proach to our plant in either regard is to be found, even in 

 halophytic species. 



In summarizing we point out the following matters of general 



interest : 



pauciflosculosa 



and of unique structure. 



2. Two kinds of hairs are present, which agree with the mor- 

 phological types found in the Compositae, as defined by Vesque. 



3. The stomata correspond closely in structure to those of 

 the majority of halophytes, as described by Warming in his 

 Halofyt Studier, but possess a more specialized inner entrance. 

 They are provided with accessory cells (Nebenzellen), which are 

 not, however, very regularly placed. 



4. The arrangement of the mesophyll possibly stands in 

 mechanical relation with the stomata. 



5. The perennial shrubby character of the plant and its pecu- 

 liar leaf anatomy separate it definitely from the genus Solidago. 



I have to thank Mr. J. E. Kirkwood for the sketch forming 

 Fig. 1. 



Bonn, March, 1901. 



