Mr. BexrnaM's. Review of the Order of Hydrophyllez. 269 
that be really the only distinction between Nemophila and Ellisia, it proves 
the expediency of uniting these two genera, more especially as in JV. parviflora 
and pedunculata the appendages are sometimes scarcely perceptible, or even 
entirely wanting, and at other times nearly half as long as the calyx. 
—— Rilisia Nyctelea has, indeed, another character, derived from the same organ, 
which distinguishes it from Nemophila, that is, the large size and expansion 
which it acquires after the fall of the corolla; but this, again, is but a charac- 
ter of degree, in respect of which the Ellisia chrysanthemifolia would stand 
intermediate between the two genera, and is not, therefore, available to sepa- 
rate them. 
The corolla of Hydrophyllec varies in general form from campanulate to 
rotate, showing but rarely (Eutoca phacelioides) an approach to the infundi- 
buliform corolla of most Borraginee. But these variations are slight, difficult 
to characterize, and so little in relation to general habit as to be unavailable 
for generic distinction. 
I have never observed in Hydrophyllece any trace of those corolline appen- 
dages, or squamae, which may be termed /aminal, from their position at the 
base of the limb of many infundibuliform corollz, whether monopetalous (as 
Borraginece), or polypetalous (as Caryophyllece); but the wnguicular or tubal 
squamee are often remarkably prominent. These squamz are analogous to 
those placed at the base of the tube of many Borraginec (having at the same 
time ligular squamz at the mouth of the tube), and to the annulus of hairs or 
scales in the tube of many Labiate. 
The /aminal squamæ are evidently of no organic importance, but mere 
excrescences of the petals, showing an approach to a transformation analo- 
gous to that by which the ligulz are conver ed into anthers, but for a different 
purpose, whatever that purpose may be. But many circumstances might in- 
duce a supposition that the case may be different with regard to the tubal 
squame. "Their origin is always below that of the stamina, and in some 
cases (as in Emmenanthe and some Eutoce) they appear to be connected with 
the filaments of the stamina in a manner analogous to the abortive filaments 
in pentandrous Caryophyllee, Paronychiee, and Amaranthacee. On the other 
hand, like the ligular squamz, although constant in form in the same species, 
they vary much in the most natural genera, such as Ajuga, Salvia and Stachys 
2N2 
