Sie ati NAO SA 
OF THE MATTO GROSSO EXPEDITION, 1891-92, 289 
province and of other Cycads further to the north-east, coupled with the entire absence 
of the order from Eastern South America, justifies us in viewing the existence of a Zamia 
at Santa Cruz as an indication of a North-Brazil-Guiana aflinity as distinguished from a 
South-Brazilian affinity in the Santa Cruz flora. 
Negative evidence is also of value in considering the nature of a flora; and although I 
do not propose to deal with this part of the subject at all exhaustively, a few facts may 
here be mentioned. I saw no species of Begonia at Santa Cruz or in the neighbourhood *. 
Now, of eighty-three species of Begonia enumerated by Alphonse de Candolle in the 
fourth volume of Martius's * Flora Brasiliensis,’ all are East Brazilian except one Amazonian 
and one from Matto Grosso. Among Malvaces the absence of the genus 4butilon—a 
genus abundantly represented in the South-Brazilian provinee—is worthy of remark, as 
is also the paucity of Composite, which in the same province appear in great force, both 
generically and specifically. Neither is the rarity of Labiatew without significance, of 
which order several genera and many species are endemic in Eastern Brazil: not one of 
these genera, such as Cunila, Hedeoma, Keithia, Glechon, was met with at Santa Cruz. 
Moreover my collection from Santa Cruz does not contain any representative of such 
preponderantly South-Brazilian genera of Scrophulariacez as Angelonia, Schwenkia, 
Brunfelsia, or Stemodia. The Gesneras, too, are wanting at Santa Cruz; according to 
Hanstein t, Eastern Brazil claims nearly 80 per cent. of Brazilian species, the localities of 
the remainder being doubtful. On the other hand the genus Drymonia, with head- 
quarters in Costa Rica, is endemic at Santa Cruz. 
The occurrence at Santa Cruz of the new genus Brosimopsis is an indication of the 
affinity of its flora with that of Amazonia, where several small or monotypie genera of 
Artocarpee flourish. Brosimum Gaudichaudii, Tréc., however, a characteristic East- 
Brazilian plant, is also met with at Santa Cruz. Unfortunately most of the orchids there 
were out of flower during our stay, but the short list of those in flower is very suggestive 
of the flora’s northern affinity. Thus the Amazonian and Guiana Cattleya superba, 
Schomb., flourishes at Santa Cruz, as also do Epidendrum imatophyllum, Lindl., a Guiana 
species, Notylia bisepala, 8. Moore, of which N. Huegelii, Reichb. f., from Mexico, seems 
the nearest ally, and Dichea cornuta, S. Moore, a species coming nearest to the Guiana 
D. graminoides, Lindl. A curious narrow-leaved Vanilla, which, when its flowers are 
known, will probably prove identical with V. ensifolia, Rolfe, from New Granada, should 
also be noticed. Lastly we have two remarkable grasses: one of these is the lowly 
Luziola pusilla, S. Moore, a species found also in Guiana; the other, a new species of 
Pariana belonging to the section with scapose inflorescences, a section found in Amazonia, 
but banished from Eastern Brazil. 
In addition to those above-named, the following is a short list of some remarkable 
Amazonian plants, or plants with Amazonian affinity, met with at Santa Cruz and in 
the neighbourhood :— 
* In fact, no species of this large tropical genus was seen by me in Matto Grosso, 
t In Martius's * Flora Brasiliensis, vol. viii. 
SECOND SERIES.— BOTANY, VOL. IV. 2Q 
