302 MR. SPENCER LE M. MOORE—PHANEROGAMIC BOTANY 
The pollen of Hornschuchia takes the form of large multicellular masses; that of 
Stormia, though often consisting of single grains, is frequently found, even when 
obtained from anthers apparently adult, to be more or less aggregated owing to per- 
sistence of the special anther-cell’s walls (Plate XXXVIII. fig. 12). Except for the 
number of its carpels, the gyneecium of Hornschuchia well stands comparison with that of 
Stormia, and the septation of the berry is another point of agreement. Bearing all these 
facts in mind, I certainly think the above-stated view of Bentham and Hooker to be that 
one for which there seems to be the most warrant. 
In the structure of its stem (Plate XX XVIII. fig. 5) Stormia presents no point of 
special interest. There is a relatively reduced pith, of which the cells are filled with 
small spheroidal or ellipsoidal starch-grains, sometimes simple, sometimes compound. In 
the xylem there are numerous (about twenty) masses of protoxylem, and tracheides are 
fairly abundant. Fibres of normal appearance make up most of the remaining part of 
the xylem, parenchyme-cells being rare. In the phloem the fibres, parenchyme, and small 
sieve-tubes with companion-cells are in every respect normal. The sections made by me 
showed usually four groups of bast-fibres; of these the three inner have their long axes 
tangential, the outer group being elongated in a radial plane. There is a well-marked 
endoderm. The medullary rays, one, two, or three cells thick in the xylem, become 
widened on passing into the phloem, and are here sometimes strengthened by single or 
aggregated sclerotic parenchyme elements; similar elements are also to be seen in both 
cortex and pith. The phellogen layer lies close to the epiderm. 
[The venation of the leaf is peculiar and reminds one more of a Clusia than of an 
Anonacea. The peculiarity is mainly owing to the presence of three nerves of tertiary 
order in the interspace between a secondary nerve and its successor. These nerves run 
more or less parallel with the secondary nerves, and they may anastomose with these or 
with each other. 'There are no stomata on the upper surface of the leaf, and the 
epidermal cells here contain each a single spheraphidal mass. The palisade parenchyme 
is one cell deep, the layer immediately below it having it cellss somewhat elongated 
perpendicularly. But few air-spaces occur in the spongy mesophyll, and even these are 
small. Immersed in the lower part of this tissue are numerous glands of normal. 
appearance. The cells of the lower epiderm usually contain one small mass of 
spheraphides; rarely two such masses are seen, or a prismatic crystal takes the place 
of the spheraphides. Stomates are numerous in this layer, each stomate having a couple 
of subsidiary cells (fig. 6). 
The arrangement of the leaf-traces is simple. The three bundles of the petiole unite at 
the axil into a single strong common bundle which courses down through two internodes, 
- and forks at the second node below that at which it entered the stem. Each of these 
forks is markedly thinner than the parent bundle, and the shorter of them inserts 
itself on the neighbouring trace of its own side close to the node, while the longer runs. 
some distance down the third internode to finally unite with the same trace. 
By referring to figs. 9-12 of Plate XXXVIII., the reader will at once understand the 
structure of the anther. As seen in transverse section, the greater part of this organ consists 
of the relatively large connective, composed mainly of a plexus of delicate cells with brown 
